Distribution of the Phenotypic Effects of Random Homologous Recombination between Two Virus Species

Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of phenotypic effects caused by genome-wide random homologous recombination has never been formally investigated. Previous data on the subject ha...

Full description

Saved in:
Bibliographic Details
Published in:PLoS pathogens Vol. 7; no. 5; p. e1002028
Main Authors: Vuillaume, Florence, Thébaud, Gaël, Urbino, Cica, Forfert, Nadège, Granier, Martine, Froissart, Rémy, Blanc, Stéphane, Peterschmitt, Michel
Format: Journal Article
Language:English
Published: United States Public Library of Science 01.05.2011
Public Library of Science (PLoS)
Subjects:
ISSN:1553-7374, 1553-7366, 1553-7374
Online Access:Get full text
Tags: Add Tag
No Tags, Be the first to tag this record!
Abstract Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of phenotypic effects caused by genome-wide random homologous recombination has never been formally investigated. Previous data on the subject have promoted the implicit view that most viral recombinant genomes are likely to be deleterious or lethal if the nucleotide identity of parental sequences is below 90%. We decided to challenge this view by creating a bank of near-random recombinants between two viral species of the genus Begomovirus (Family Geminiviridae) exhibiting 82% nucleotide identity, and by testing infectivity and in planta accumulation of recombinant clones randomly extracted from this bank. The bank was created by DNA-shuffling-a technology initially applied to the random shuffling of individual genes, and here implemented for the first time to shuffle full-length viral genomes. Together with our previously described system allowing the direct cloning of full-length infectious geminivirus genomes, it provided a unique opportunity to generate hundreds of "mosaic" virus genomes, directly testable for infectivity. A subset of 47 randomly chosen recombinants was sequenced, individually inoculated into tomato plants, and compared with the parental viruses. Surprisingly, our results showed that all recombinants were infectious and accumulated at levels comparable or intermediate to that of the parental clones. This indicates that, in our experimental system, despite the fact that the parental genomes differ by nearly 20%, lethal and/or large deleterious effects of recombination are very rare, in striking contrast to the common view that has emerged from previous studies published on other viruses.
AbstractList   Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of phenotypic effects caused by genome-wide random homologous recombination has never been formally investigated. Previous data on the subject have promoted the implicit view that most viral recombinant genomes are likely to be deleterious or lethal if the nucleotide identity of parental sequences is below 90%. We decided to challenge this view by creating a bank of near-random recombinants between two viral species of the genus Begomovirus (Family Geminiviridae) exhibiting 82% nucleotide identity, and by testing infectivity and in planta accumulation of recombinant clones randomly extracted from this bank. The bank was created by DNA-shuffling--a technology initially applied to the random shuffling of individual genes, and here implemented for the first time to shuffle full-length viral genomes. Together with our previously described system allowing the direct cloning of full-length infectious geminivirus genomes, it provided a unique opportunity to generate hundreds of "mosaic" virus genomes, directly testable for infectivity. A subset of 47 randomly chosen recombinants was sequenced, individually inoculated into tomato plants, and compared with the parental viruses. Surprisingly, our results showed that all recombinants were infectious and accumulated at levels comparable or intermediate to that of the parental clones. This indicates that, in our experimental system, despite the fact that the parental genomes differ by nearly 20%, lethal and/or large deleterious effects of recombination are very rare, in striking contrast to the common view that has emerged from previous studies published on other viruses.
Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of phenotypic effects caused by genome-wide random homologous recombination has never been formally investigated. Previous data on the subject have promoted the implicit view that most viral recombinant genomes are likely to be deleterious or lethal if the nucleotide identity of parental sequences is below 90%. We decided to challenge this view by creating a bank of near-random recombinants between two viral species of the genus Begomovirus (Family Geminiviridae) exhibiting 82% nucleotide identity, and by testing infectivity and in planta accumulation of recombinant clones randomly extracted from this bank. The bank was created by DNA-shuffling-a technology initially applied to the random shuffling of individual genes, and here implemented for the first time to shuffle full-length viral genomes. Together with our previously described system allowing the direct cloning of full-length infectious geminivirus genomes, it provided a unique opportunity to generate hundreds of "mosaic" virus genomes, directly testable for infectivity. A subset of 47 randomly chosen recombinants was sequenced, individually inoculated into tomato plants, and compared with the parental viruses. Surprisingly, our results showed that all recombinants were infectious and accumulated at levels comparable or intermediate to that of the parental clones. This indicates that, in our experimental system, despite the fact that the parental genomes differ by nearly 20%, lethal and/or large deleterious effects of recombination are very rare, in striking contrast to the common view that has emerged from previous studies published on other viruses.
Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of phenotypic effects caused by genome-wide random homologous recombination has never been formally investigated. Previous data on the subject have promoted the implicit view that most viral recombinant genomes are likely to be deleterious or lethal if the nucleotide identity of parental sequences is below 90%. We decided to challenge this view by creating a bank of near-random recombinants between two viral species of the genus Begomovirus (Family Geminiviridae) exhibiting 82% nucleotide identity, and by testing infectivity and in planta accumulation of recombinant clones randomly extracted from this bank. The bank was created by DNA-shuffling-a technology initially applied to the random shuffling of individual genes, and here implemented for the first time to shuffle full-length viral genomes. Together with our previously described system allowing the direct cloning of full-length infectious geminivirus genomes, it provided a unique opportunity to generate hundreds of "mosaic" virus genomes, directly testable for infectivity. A subset of 47 randomly chosen recombinants was sequenced, individually inoculated into tomato plants, and compared with the parental viruses. Surprisingly, our results showed that all recombinants were infectious and accumulated at levels comparable or intermediate to that of the parental clones. This indicates that, in our experimental system, despite the fact that the parental genomes differ by nearly 20%, lethal and/or large deleterious effects of recombination are very rare, in striking contrast to the common view that has emerged from previous studies published on other viruses.Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of phenotypic effects caused by genome-wide random homologous recombination has never been formally investigated. Previous data on the subject have promoted the implicit view that most viral recombinant genomes are likely to be deleterious or lethal if the nucleotide identity of parental sequences is below 90%. We decided to challenge this view by creating a bank of near-random recombinants between two viral species of the genus Begomovirus (Family Geminiviridae) exhibiting 82% nucleotide identity, and by testing infectivity and in planta accumulation of recombinant clones randomly extracted from this bank. The bank was created by DNA-shuffling-a technology initially applied to the random shuffling of individual genes, and here implemented for the first time to shuffle full-length viral genomes. Together with our previously described system allowing the direct cloning of full-length infectious geminivirus genomes, it provided a unique opportunity to generate hundreds of "mosaic" virus genomes, directly testable for infectivity. A subset of 47 randomly chosen recombinants was sequenced, individually inoculated into tomato plants, and compared with the parental viruses. Surprisingly, our results showed that all recombinants were infectious and accumulated at levels comparable or intermediate to that of the parental clones. This indicates that, in our experimental system, despite the fact that the parental genomes differ by nearly 20%, lethal and/or large deleterious effects of recombination are very rare, in striking contrast to the common view that has emerged from previous studies published on other viruses.
Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of phenotypic effects caused by genome-wide random homologous recombination has never been formally investigated. Previous data on the subject have promoted the implicit view that most viral recombinant genomes are likely to be deleterious or lethal if the nucleotide identity of parental sequences is below 90%. We decided to challenge this view by creating a bank of near-random recombinants between two viral species of the genus Begomovirus (Family Geminiviridae) exhibiting 82% nucleotide identity, and by testing infectivity and in planta accumulation of recombinant clones randomly extracted from this bank. The bank was created by DNA-shuffling-a technology initially applied to the random shuffling of individual genes, and here implemented for the first time to shuffle full-length viral genomes. Together with our previously described system allowing the direct cloning of full-length infectious geminivirus genomes, it provided a unique opportunity to generate hundreds of "mosaic" virus genomes, directly testable for infectivity. A subset of 47 randomly chosen recombinants was sequenced, individually inoculated into tomato plants, and compared with the parental viruses. Surprisingly, our results showed that all recombinants were infectious and accumulated at levels comparable or intermediate to that of the parental clones. This indicates that, in our experimental system, despite the fact that the parental genomes differ by nearly 20%, lethal and/or large deleterious effects of recombination are very rare, in striking contrast to the common view that has emerged from previous studies published on other viruses. Recombination creates new genome combinations by joining genome fragments of distinct "parental" origin. This phenomenon, frequent in viral populations, combines mutations originally present on distinct parental genomes, increasing genetic diversity and creating "offspring" with altered biological properties. Consistently, recombination is often associated with the emergence of economically important viruses, with modified host range and virulence. In fact, recombination events can be lethal, deleterious or beneficial, but the respective frequency of these phenotypic effects is unknown and unpredictable. A generally accepted view, which we formally challenge in the present paper, is that most viral recombination events are deleterious or lethal when the parental sequences diverge by more than 10%. However, at present, no dedicated data set supports this supposition. We generated hundreds of "mosaic" genomes randomly from two plant virus species diverging by 18%, and tested a subset of 47 of these recombinants for viability and within-host accumulation. Surprisingly, all were viable, replicated, and accumulated at a pace comparable to that of the parents. Our results are in striking contrast to the current view, and show that viral recombination can have little phenotypic effect, at least in some cases, even when the parental sequences diverge by far more than 10%.
Author Summary : Recombination creates new genome combinations by joining genome fragments of distinct “parental” origin. This phenomenon, frequent in viral populations, combines mutations originally present on distinct parental genomes, increasing genetic diversity and creating “offspring” with altered biological properties. Consistently, recombination is often associated with the emergence of economically important viruses, with modified host range and virulence. In fact, recombination events can be lethal, deleterious or beneficial, but the respective frequency of these phenotypic effects is unknown and unpredictable. A generally accepted view, which we formally challenge in the present paper, is that most viral recombination events are deleterious or lethal when the parental sequences diverge by more than 10%. However, at present, no dedicated data set supports this supposition. We generated hundreds of “mosaic” genomes randomly from two plant virus species diverging by 18%, and tested a subset of 47 of these recombinants for viability and within-host accumulation. Surprisingly, all were viable, replicated, and accumulated at a pace comparable to that of the parents. Our results are in striking contrast to the current view, and show that viral recombination can have little phenotypic effect, at least in some cases, even when the parental sequences diverge by far more than 10%.
Audience Academic
Author Urbino, Cica
Peterschmitt, Michel
Froissart, Rémy
Vuillaume, Florence
Forfert, Nadège
Thébaud, Gaël
Blanc, Stéphane
Granier, Martine
AuthorAffiliation 2 Laboratoire Maladies Infectieuses & Vecteurs: Écologie, Génétique, Évolution & Contrôle (MIVEGEC), CNRS-IRD-Université de Montpellier I, Agropolis, Montpellier, France
1 CIRAD, INRA, CNRS – Unité mixte de recherche Biologie & génétique des interactions plante-parasite (BGPI), Montpellier, France
Cornell University, United States of America
AuthorAffiliation_xml – name: Cornell University, United States of America
– name: 1 CIRAD, INRA, CNRS – Unité mixte de recherche Biologie & génétique des interactions plante-parasite (BGPI), Montpellier, France
– name: 2 Laboratoire Maladies Infectieuses & Vecteurs: Écologie, Génétique, Évolution & Contrôle (MIVEGEC), CNRS-IRD-Université de Montpellier I, Agropolis, Montpellier, France
Author_xml – sequence: 1
  givenname: Florence
  surname: Vuillaume
  fullname: Vuillaume, Florence
– sequence: 2
  givenname: Gaël
  surname: Thébaud
  fullname: Thébaud, Gaël
– sequence: 3
  givenname: Cica
  surname: Urbino
  fullname: Urbino, Cica
– sequence: 4
  givenname: Nadège
  surname: Forfert
  fullname: Forfert, Nadège
– sequence: 5
  givenname: Martine
  surname: Granier
  fullname: Granier, Martine
– sequence: 6
  givenname: Rémy
  surname: Froissart
  fullname: Froissart, Rémy
– sequence: 7
  givenname: Stéphane
  surname: Blanc
  fullname: Blanc, Stéphane
– sequence: 8
  givenname: Michel
  surname: Peterschmitt
  fullname: Peterschmitt, Michel
BackLink https://www.ncbi.nlm.nih.gov/pubmed/21573141$$D View this record in MEDLINE/PubMed
https://hal.inrae.fr/hal-02647631$$DView record in HAL
BookMark eNqVk99v0zAQxyM0xH7Af4AgEg9oDy3-kTjJHpCqMWilClA3eLVs59J6SuwQOxv773HbbFqrCYTyEOvu873zne-OowNjDUTRa4zGmGb4w7XtOyPqcdsKP8YIEUTyZ9ERTlM6ymiWHDw6H0bHzl0jlGCK2YvokOA0ozjBR5H6pJ3vtOy9tia2VexXEH9fgbH-rtUqvqgqUN6tPQthStvEU9vY2i5t7-IFKNtIbcRGLMHfApj46tbGP3UX_JctKA3uZfS8ErWDV8P_JPrx-eLqfDqaf_syO5_MRyojzI8UkoliBAqksMxJlctQUThIyiTBIq8oEyWWSgJQlBcSUpJlSBGUMkErnNCT6O02bltbx4f-OI5JXmCEU0YCMdsSpRXXvO10I7o7boXmG4Ptllx0XqsaOMvSHCTkUhGWMIxEkhKqgIkcSkZlEWJ9HLL1soFSgfGdqHeC7nqMXvGlveHh8nlGaAhwug2w2pNNJ3O-tqGQOWMU3-DAvh-SdfZXD87zRjsFdS0MhJfgBSKUEZTQf5I5yyhJkhQF8t0e-XTLBmopQle0qWwoRq1j8glJA8ZwkQVq_AQVvhIarcLgVjrYdwSnO4LAePjtl6J3js8uF__Bft1l3zx-lYe-3k98AJItoDrrXAfVA4IRXy_WfRf4erH4sFhBdrYnU9pv5j4Uquu_i_8AfIIoeg
CitedBy_id crossref_primary_10_1111_eva_12157
crossref_primary_10_3390_agronomy15051006
crossref_primary_10_1016_j_coviro_2014_12_005
crossref_primary_10_1007_s13595_015_0487_4
crossref_primary_10_1371_journal_pone_0174651
crossref_primary_10_1099_jgv_0_000633
crossref_primary_10_1016_j_virol_2020_01_002
crossref_primary_10_3390_v3091699
crossref_primary_10_1016_j_coviro_2018_08_013
crossref_primary_10_24072_pcjournal_379
crossref_primary_10_1186_1743_422X_11_66
crossref_primary_10_7717_peerj_1320
crossref_primary_10_1016_j_virol_2015_06_019
crossref_primary_10_1038_srep08153
crossref_primary_10_1128_JVI_00541_21
crossref_primary_10_1371_journal_pone_0058375
Cites_doi 10.1099/vir.0.83077-0
10.1073/pnas.91.22.10747
10.1128/JVI.02152-08
10.1016/j.virol.2006.02.028
10.1099/0022-1317-81-7-1839
10.1128/JVI.74.9.4214-4219.2000
10.1099/vir.0.18894-0
10.1371/journal.ppat.0030181
10.1016/j.jviromet.2007.09.015
10.1146/annurev.genet.36.040202.111115
10.1186/1743-422X-5-61
10.1371/journal.pgen.0010051
10.1063/1.3449300
10.1110/ps.0306603
10.1186/1743-422X-5-135
10.1016/j.virol.2003.10.002
10.1006/viro.2002.1633
10.1094/PDIS.1999.83.3.303B
10.1016/j.virol.2004.11.017
10.1093/nar/29.9.e45
10.1128/JVI.00524-07
10.1534/genetics.105.051136
10.1128/JVI.00337-08
10.1094/PDIS.1999.83.11.1006
10.1099/0022-1317-78-8-2101
10.1098/rstb.2010.0068
10.1073/pnas.0500729102
10.1099/vir.0.83252-0
10.1038/nrg2323
10.1073/pnas.0400146101
10.1016/j.meegid.2008.07.010
10.1016/0042-6822(91)90795-D
10.1371/journal.pcbi.1000178
10.1099/vir.0.80805-0
10.1371/journal.pbio.0030089
10.1128/JVI.79.9.5732-5742.2005
10.1006/viro.1999.0056
10.1371/journal.ppat.0030008
ContentType Journal Article
Copyright COPYRIGHT 2011 Public Library of Science
2011 Vuillaume et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited: Vuillaume F, Thébaud G, Urbino C, Forfert N, Granier M, et al. (2011) Distribution of the Phenotypic Effects of Random Homologous Recombination between Two Virus Species. PLoS Pathog 7(5): e1002028. doi:10.1371/journal.ppat.1002028
Attribution
Vuillaume et al. 2011
Copyright_xml – notice: COPYRIGHT 2011 Public Library of Science
– notice: 2011 Vuillaume et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited: Vuillaume F, Thébaud G, Urbino C, Forfert N, Granier M, et al. (2011) Distribution of the Phenotypic Effects of Random Homologous Recombination between Two Virus Species. PLoS Pathog 7(5): e1002028. doi:10.1371/journal.ppat.1002028
– notice: Attribution
– notice: Vuillaume et al. 2011
DBID AAYXX
CITATION
CGR
CUY
CVF
ECM
EIF
NPM
ISN
ISR
3V.
7QL
7U9
7X7
7XB
88E
8FE
8FH
8FI
8FJ
8FK
ABUWG
AEUYN
AFKRA
AZQEC
BBNVY
BENPR
BHPHI
C1K
CCPQU
COVID
DWQXO
FYUFA
GHDGH
GNUQQ
H94
HCIFZ
K9.
LK8
M0S
M1P
M7P
PHGZM
PHGZT
PIMPY
PJZUB
PKEHL
PPXIY
PQEST
PQGLB
PQQKQ
PQUKI
PRINS
7X8
1XC
VOOES
5PM
DOA
DOI 10.1371/journal.ppat.1002028
DatabaseName CrossRef
Medline
MEDLINE
MEDLINE (Ovid)
MEDLINE
MEDLINE
PubMed
Gale In Context: Canada
Gale In Context: Science
ProQuest Central (Corporate)
Bacteriology Abstracts (Microbiology B)
Virology and AIDS Abstracts
Health & Medical Collection
ProQuest Central (purchase pre-March 2016)
Medical Database (Alumni Edition)
ProQuest SciTech Collection
ProQuest Natural Science Collection
ProQuest Hospital Collection
Hospital Premium Collection (Alumni Edition)
ProQuest Central (Alumni) (purchase pre-March 2016)
ProQuest Central (Alumni)
ProQuest One Sustainability
ProQuest Central UK/Ireland
ProQuest Central Essentials
Biological Science Collection
ProQuest Central
Natural Science Collection
Environmental Sciences and Pollution Management
ProQuest One
Coronavirus Research Database
ProQuest Central
Health Research Premium Collection
Health Research Premium Collection (Alumni)
ProQuest Central Student
AIDS and Cancer Research Abstracts
SciTech Premium Collection
ProQuest Health & Medical Complete (Alumni)
Biological Sciences
ProQuest Health & Medical Collection
Medical Database
Biological Science Database
ProQuest Central Premium
ProQuest One Academic (New)
Publicly Available Content Database
ProQuest Health & Medical Research Collection
ProQuest One Academic Middle East (New)
ProQuest One Health & Nursing
ProQuest One Academic Eastern Edition (DO NOT USE)
ProQuest One Applied & Life Sciences
ProQuest One Academic (retired)
ProQuest One Academic UKI Edition
ProQuest Central China
MEDLINE - Academic
Hyper Article en Ligne (HAL)
Hyper Article en Ligne (HAL) (Open Access)
PubMed Central (Full Participant titles)
DOAJ Open Access Full Text
DatabaseTitle CrossRef
MEDLINE
Medline Complete
MEDLINE with Full Text
PubMed
MEDLINE (Ovid)
Publicly Available Content Database
ProQuest Central Student
ProQuest One Academic Middle East (New)
ProQuest Central Essentials
ProQuest Health & Medical Complete (Alumni)
ProQuest Central (Alumni Edition)
SciTech Premium Collection
ProQuest One Community College
ProQuest One Health & Nursing
ProQuest Natural Science Collection
ProQuest Central China
Environmental Sciences and Pollution Management
ProQuest Central
ProQuest One Applied & Life Sciences
ProQuest One Sustainability
ProQuest Health & Medical Research Collection
Health Research Premium Collection
Health and Medicine Complete (Alumni Edition)
Natural Science Collection
ProQuest Central Korea
Bacteriology Abstracts (Microbiology B)
Health & Medical Research Collection
Biological Science Collection
AIDS and Cancer Research Abstracts
ProQuest Central (New)
ProQuest Medical Library (Alumni)
Virology and AIDS Abstracts
ProQuest Biological Science Collection
ProQuest One Academic Eastern Edition
Coronavirus Research Database
ProQuest Hospital Collection
Health Research Premium Collection (Alumni)
Biological Science Database
ProQuest SciTech Collection
ProQuest Hospital Collection (Alumni)
ProQuest Health & Medical Complete
ProQuest Medical Library
ProQuest One Academic UKI Edition
ProQuest One Academic
ProQuest One Academic (New)
ProQuest Central (Alumni)
MEDLINE - Academic
DatabaseTitleList
MEDLINE

MEDLINE - Academic
AIDS and Cancer Research Abstracts
Publicly Available Content Database



Database_xml – sequence: 1
  dbid: DOA
  name: DOAJ Directory of Open Access Journals
  url: https://www.doaj.org/
  sourceTypes: Open Website
– sequence: 2
  dbid: NPM
  name: PubMed
  url: http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?db=PubMed
  sourceTypes: Index Database
– sequence: 3
  dbid: PIMPY
  name: Publicly Available Content Database
  url: http://search.proquest.com/publiccontent
  sourceTypes: Aggregation Database
DeliveryMethod fulltext_linktorsrc
Discipline Biology
DocumentTitleAlternate Phenotypic Effects of Random Recombination
EISSN 1553-7374
ExternalDocumentID 1289101562
oai_doaj_org_article_6758ebe8bc264610a4523ce6a8ed63b9
PMC3088723
oai:HAL:hal-02647631v1
2896597821
A258916197
21573141
10_1371_journal_ppat_1002028
Genre Research Support, Non-U.S. Gov't
Journal Article
GeographicLocations France
GeographicLocations_xml – name: France
GroupedDBID ---
123
29O
2WC
53G
5VS
7X7
88E
8FE
8FH
8FI
8FJ
AAFWJ
AAUCC
AAWOE
AAYXX
ABDBF
ABUWG
ACCTH
ACGFO
ACIHN
ACPRK
ACUHS
ADBBV
ADRAZ
AEAQA
AENEX
AEUYN
AFFHD
AFKRA
AFPKN
AFRAH
AHMBA
ALMA_UNASSIGNED_HOLDINGS
AOIJS
B0M
BAWUL
BBNVY
BCNDV
BENPR
BHPHI
BPHCQ
BVXVI
BWKFM
CCPQU
CITATION
CS3
DIK
DU5
E3Z
EAP
EAS
EBD
EMK
EMOBN
ESX
F5P
FPL
FYUFA
GROUPED_DOAJ
GX1
HCIFZ
HMCUK
HYE
IAO
IHR
INH
INR
ISN
ISR
ITC
KQ8
LK8
M1P
M48
M7P
MM.
O5R
O5S
OK1
OVT
P2P
PGMZT
PHGZM
PHGZT
PIMPY
PJZUB
PPXIY
PQGLB
PQQKQ
PROAC
PSQYO
QN7
RNS
RPM
SV3
TR2
TUS
UKHRP
WOW
~8M
3V.
CGR
CUY
CVF
ECM
EIF
H13
IPNFZ
M~E
NPM
PV9
QF4
RIG
RZL
WOQ
7QL
7U9
7XB
8FK
AZQEC
C1K
COVID
DWQXO
GNUQQ
H94
K9.
PKEHL
PQEST
PQUKI
PRINS
7X8
PUEGO
1XC
VOOES
5PM
AAPBV
ABPTK
ID FETCH-LOGICAL-c726t-c0b4c62e90c1b82f8b028b82b36b21a8f36ad1bcbee3089be52770c2056a3f143
IEDL.DBID DOA
ISICitedReferencesCount 18
ISICitedReferencesURI http://www.webofscience.com/api/gateway?GWVersion=2&SrcApp=Summon&SrcAuth=ProQuest&DestLinkType=CitingArticles&DestApp=WOS_CPL&KeyUT=000291014000014&url=https%3A%2F%2Fcvtisr.summon.serialssolutions.com%2F%23%21%2Fsearch%3Fho%3Df%26include.ft.matches%3Dt%26l%3Dnull%26q%3D
ISSN 1553-7374
1553-7366
IngestDate Sun Oct 01 00:11:19 EDT 2023
Tue Oct 14 19:06:43 EDT 2025
Tue Nov 04 01:57:57 EST 2025
Tue Oct 14 20:43:05 EDT 2025
Fri Sep 05 10:32:01 EDT 2025
Thu Oct 02 09:40:28 EDT 2025
Sat Nov 29 14:37:51 EST 2025
Tue Nov 11 10:30:01 EST 2025
Tue Nov 04 17:57:55 EST 2025
Thu Nov 13 14:47:33 EST 2025
Thu Nov 13 14:22:39 EST 2025
Wed Feb 19 02:05:04 EST 2025
Sat Nov 29 04:10:11 EST 2025
Tue Nov 18 22:28:03 EST 2025
IsDoiOpenAccess true
IsOpenAccess true
IsPeerReviewed true
IsScholarly true
Issue 5
Keywords VIROLOGIE
BEGOMOVIRUS
Language English
License Attribution: http://creativecommons.org/licenses/by
This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are properly credited.
Creative Commons Attribution License
LinkModel DirectLink
MergedId FETCHMERGED-LOGICAL-c726t-c0b4c62e90c1b82f8b028b82b36b21a8f36ad1bcbee3089be52770c2056a3f143
Notes ObjectType-Article-1
SourceType-Scholarly Journals-1
ObjectType-Feature-2
content type line 14
content type line 23
ObjectType-Article-2
ObjectType-Feature-1
Conceived and designed the experiments: SB MP. Performed the experiments: FV CU NF MG MP. Analyzed the data: FV GT SB MP. Wrote the paper: FV GT RF SB MP.
ORCID 0000-0002-3412-0989
0000-0001-8234-1308
0000-0002-2987-4997
0000-0002-6215-9389
OpenAccessLink https://doaj.org/article/6758ebe8bc264610a4523ce6a8ed63b9
PMID 21573141
PQID 1289101562
PQPubID 1436335
ParticipantIDs plos_journals_1289101562
doaj_primary_oai_doaj_org_article_6758ebe8bc264610a4523ce6a8ed63b9
pubmedcentral_primary_oai_pubmedcentral_nih_gov_3088723
hal_primary_oai_HAL_hal_02647631v1
proquest_miscellaneous_902362043
proquest_miscellaneous_867324450
proquest_journals_1289101562
gale_infotracmisc_A258916197
gale_infotracacademiconefile_A258916197
gale_incontextgauss_ISR_A258916197
gale_incontextgauss_ISN_A258916197
pubmed_primary_21573141
crossref_primary_10_1371_journal_ppat_1002028
crossref_citationtrail_10_1371_journal_ppat_1002028
PublicationCentury 2000
PublicationDate 2011-05-01
PublicationDateYYYYMMDD 2011-05-01
PublicationDate_xml – month: 05
  year: 2011
  text: 2011-05-01
  day: 01
PublicationDecade 2010
PublicationPlace United States
PublicationPlace_xml – name: United States
– name: San Francisco
– name: San Francisco, USA
PublicationTitle PLoS pathogens
PublicationTitleAlternate PLoS Pathog
PublicationYear 2011
Publisher Public Library of Science
Public Library of Science (PLoS)
Publisher_xml – name: Public Library of Science
– name: Public Library of Science (PLoS)
References R Froissart (ref34) 2010; 365
WPC Stemmer (ref27) 1994; 91
(ref39) 2010
S Wain-Hobson (ref5) 2003; 84
C Urbino (ref35) 2008; 5
S De Rozieres (ref9) 2008; 82
A Bruyere (ref14) 2000; 74
J Bonnet (ref13) 2005; 332
MW Pfaffl (ref38) 2001; 29
LR Banner (ref20) 1991; 185
S Duffy (ref1) 2008; 9
D Posada (ref19) 2002; 36
DA Drummond (ref6) 2005; 102
P Carrasco (ref8) 2007; 81
IM Moreno (ref16) 2004; 318
B Pico (ref36) 1999; 83
X Zhou (ref26) 1997; 78
O Pierrugues (ref12) 2007; 88
M Padidam (ref22) 1999; 265
P Lefeuvre (ref28) 2007; 88
F Escriu (ref15) 2007; 3
DP Martin (ref10) 2005; 1
P Agudelo-Romero (ref29) 2008; 8
P Lefeuvre (ref21) 2007; 3
P Lefeuvre (ref18) 2009; 83
SF Elena (ref2) 2010; 20
S Duffy (ref30) 2006; 172
G Mason (ref37) 2008; 147
A Urbanowicz (ref4) 2005; 79
F Monci (ref24) 2002; 303
AI Sanz (ref25) 2000; 81
MM Meyer (ref31) 2003; 12
M Peterschmitt (ref33) 1999; 83
H Delatte (ref32) 2005; 86
R Sanjuan (ref7) 2004; 101
J Archer (ref17) 2008; 4
S Garcia-Andres (ref23) 2006; 350
R Froissart (ref3) 2005; 3
E van der Walt (ref11) 2008; 5
References_xml – volume: 88
  start-page: 2852
  year: 2007
  ident: ref12
  article-title: Biological properties and relative fitness of inter-subgroup Cucumber mosaic virus RNA 3 recombinants produced in vitro.
  publication-title: J Gen Virol
  doi: 10.1099/vir.0.83077-0
– volume: 91
  start-page: 10747
  year: 1994
  ident: ref27
  article-title: DNA Shuffling by Random Fragmentation and Reassembly - in-Vitro Recombination for Molecular Evolution.
  publication-title: Proc Natl Acad Sci U S A
  doi: 10.1073/pnas.91.22.10747
– volume: 83
  start-page: 2697
  year: 2009
  ident: ref18
  article-title: Widely Conserved Recombination Patterns among Single-Stranded DNA Viruses.
  publication-title: J Virol
  doi: 10.1128/JVI.02152-08
– volume: 350
  start-page: 433
  year: 2006
  ident: ref23
  article-title: Begomovirus genetic diversity in the native plant reservoir Solanum nigrum: Evidence for the presence of a new virus species of recombinant nature.
  publication-title: Virology
  doi: 10.1016/j.virol.2006.02.028
– volume: 81
  start-page: 1839
  year: 2000
  ident: ref25
  article-title: Multiple infection, recombination and genome relationships among begomovirus isolates found in cotton and other plants in Pakistan.
  publication-title: J Gen Virol
  doi: 10.1099/0022-1317-81-7-1839
– volume: 74
  start-page: 4214
  year: 2000
  ident: ref14
  article-title: Frequent homologous recombination events between molecules of one RNA component in a multipartite RNA virus.
  publication-title: J Virol
  doi: 10.1128/JVI.74.9.4214-4219.2000
– volume: 84
  start-page: 885
  year: 2003
  ident: ref5
  article-title: Network analysis of human and simian immunodeficiency virus sequence sets reveals massive recombination resulting in shorter pathways.
  publication-title: J Gen Virol
  doi: 10.1099/vir.0.18894-0
– volume: 3
  start-page: 1782
  year: 2007
  ident: ref21
  article-title: Avoidance of protein fold disruption in natural virus recombinants.
  publication-title: Plos Pathog
  doi: 10.1371/journal.ppat.0030181
– volume: 147
  start-page: 282
  year: 2008
  ident: ref37
  article-title: Real-time PCR for the quantitation of Tomato yellow leaf curl Sardinia virus in tomato plants and in Bemisia tabaci.
  publication-title: J Virol Methods
  doi: 10.1016/j.jviromet.2007.09.015
– volume: 36
  start-page: 75
  year: 2002
  ident: ref19
  article-title: Recombination in evolutionary genomics.
  publication-title: Annu Rev Genet
  doi: 10.1146/annurev.genet.36.040202.111115
– volume: 5
  start-page: 11
  year: 2008
  ident: ref11
  article-title: Viable chimaeric viruses confirm the biological importance of sequence specific Maize streak virus movement protein and coat protein interactions.
  publication-title: Virol J
  doi: 10.1186/1743-422X-5-61
– volume: 1
  start-page: 475
  year: 2005
  ident: ref10
  article-title: The evolutionary value of recombination is constrained by genome modularity.
  publication-title: Plos Genet
  doi: 10.1371/journal.pgen.0010051
– year: 2010
  ident: ref39
  article-title: R: A Language and Environment for Statistical Computing.
– volume: 20
  start-page: 12
  year: 2010
  ident: ref2
  article-title: Simple genomes, complex interactions: Epistasis in RNA virus.
  publication-title: Chaos
  doi: 10.1063/1.3449300
– volume: 12
  start-page: 1686
  year: 2003
  ident: ref31
  article-title: Library analysis of SCHEMA-guided protein recombination.
  publication-title: Protein Sci
  doi: 10.1110/ps.0306603
– volume: 5
  start-page: 10
  year: 2008
  ident: ref35
  article-title: A novel cloning strategy for isolating, genotyping and phenotyping genetic variants of geminiviruses.
  publication-title: Virol J
  doi: 10.1186/1743-422X-5-135
– volume: 318
  start-page: 451
  year: 2004
  ident: ref16
  article-title: Variability and genetic structure of the population of Watermelon mosaic virus infecting melon in Spain.
  publication-title: Virology
  doi: 10.1016/j.virol.2003.10.002
– volume: 303
  start-page: 317
  year: 2002
  ident: ref24
  article-title: A natural recombinant between the geminiviruses Tomato yellow leaf curl Sardinia virus and Tomato yellow leaf curl virus exhibits a novel pathogenic phenotype and is becoming prevalent in Spanish populations.
  publication-title: Virology
  doi: 10.1006/viro.2002.1633
– volume: 83
  start-page: 303
  year: 1999
  ident: ref33
  article-title: First report of Tomato yellow leaf curl virus in Réunion Island.
  publication-title: Plant Dis
  doi: 10.1094/PDIS.1999.83.3.303B
– volume: 332
  start-page: 359
  year: 2005
  ident: ref13
  article-title: Role of recombination in the evolution of natural populations of Cucumber mosaic virus, a tripartite RNA plant virus.
  publication-title: Virology
  doi: 10.1016/j.virol.2004.11.017
– volume: 29
  start-page: 2002
  year: 2001
  ident: ref38
  article-title: A new mathematical model for relative quantification in real-time RT-PCR.
  publication-title: Nucleic Acids Res
  doi: 10.1093/nar/29.9.e45
– volume: 81
  start-page: 12979
  year: 2007
  ident: ref8
  article-title: Distribution of fitness and virulence effects caused by single-nucleotide substitutions in Tobacco etch virus.
  publication-title: J Virol
  doi: 10.1128/JVI.00524-07
– volume: 172
  start-page: 751
  year: 2006
  ident: ref30
  article-title: Pleiotropic costs of niche expansion in the RNA bacteriophage Phi 6.
  publication-title: Genetics
  doi: 10.1534/genetics.105.051136
– volume: 82
  start-page: 7953
  year: 2008
  ident: ref9
  article-title: Replication properties of clade a/c chimeric feline immunodeficiency viruses and evaluation of infection kinetics in the domestic cat.
  publication-title: J Virol
  doi: 10.1128/JVI.00337-08
– volume: 83
  start-page: 1006
  year: 1999
  ident: ref36
  article-title: Improved diagnostic techniques for Tomato yellow leaf curl virus in tomato breeding programs.
  publication-title: Plant Dis
  doi: 10.1094/PDIS.1999.83.11.1006
– volume: 78
  start-page: 2101
  year: 1997
  ident: ref26
  article-title: Evidence that DNA-A of a geminivirus associated with severe cassava mosaic disease in Uganda has arisen by interspecific recombination.
  publication-title: J Gen Virol
  doi: 10.1099/0022-1317-78-8-2101
– volume: 365
  start-page: 1907
  year: 2010
  ident: ref34
  article-title: The virulence-transmission trade-off in vector-borne plant viruses: a review of (non-) existing studies.
  publication-title: Philos Trans R Soc B Biol Sci
  doi: 10.1098/rstb.2010.0068
– volume: 102
  start-page: 5380
  year: 2005
  ident: ref6
  article-title: On the conservative nature of intragenic recombination.
  publication-title: Proc Natl Acad Sci U S A
  doi: 10.1073/pnas.0500729102
– volume: 88
  start-page: 3458
  year: 2007
  ident: ref28
  article-title: Begomovirus ‘melting pot’ in the south-west Indian Ocean islands: molecular diversity and evolution through recombination.
  publication-title: J Gen Virol
  doi: 10.1099/vir.0.83252-0
– volume: 9
  start-page: 267
  year: 2008
  ident: ref1
  article-title: Rates of evolutionary change in viruses: patterns and determinants.
  publication-title: Nat Rev Genet
  doi: 10.1038/nrg2323
– volume: 101
  start-page: 8396
  year: 2004
  ident: ref7
  article-title: The distribution of fitness effects caused by single-nucleotide substitutions in an RNA virus.
  publication-title: Proc Natl Acad Sci U S A
  doi: 10.1073/pnas.0400146101
– volume: 8
  start-page: 806
  year: 2008
  ident: ref29
  article-title: The pleiotropic cost of host-specialization in Tobacco etch potyvirus.
  publication-title: Infection Genetics and Evolution
  doi: 10.1016/j.meegid.2008.07.010
– volume: 185
  start-page: 441
  year: 1991
  ident: ref20
  article-title: Random nature of Coronavirus RNA Recombination in the absence of selection pressure.
  publication-title: Virology
  doi: 10.1016/0042-6822(91)90795-D
– volume: 4
  start-page: 7
  year: 2008
  ident: ref17
  article-title: Identifying the important HIV-1 recombination breakpoints.
  publication-title: Plos Comput Biol
  doi: 10.1371/journal.pcbi.1000178
– volume: 86
  start-page: 1533
  year: 2005
  ident: ref32
  article-title: South West Indian Ocean islands tomato begomovirus populations represent a new major monopartite begomovirus group.
  publication-title: J Gen Virol
  doi: 10.1099/vir.0.80805-0
– volume: 3
  start-page: 389
  year: 2005
  ident: ref3
  article-title: Recombination every day: Abundant recombination in a virus during a single multi-cellular host infection.
  publication-title: Plos Biol
  doi: 10.1371/journal.pbio.0030089
– volume: 79
  start-page: 5732
  year: 2005
  ident: ref4
  article-title: Homologous crossovers among molecules of Brome mosaic bromovirus RNA1 or RNA2 segments in vivo.
  publication-title: J Virol
  doi: 10.1128/JVI.79.9.5732-5742.2005
– volume: 265
  start-page: 218
  year: 1999
  ident: ref22
  article-title: Possible emergence of new geminiviruses by frequent recombination.
  publication-title: Virology
  doi: 10.1006/viro.1999.0056
– volume: 3
  start-page: 67
  year: 2007
  ident: ref15
  article-title: Constraints to genetic exchange support gene coadaptation in a tripartite RNA virus.
  publication-title: Plos Pathog
  doi: 10.1371/journal.ppat.0030008
SSID ssj0041316
Score 2.0904553
Snippet Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of...
Author Summary : Recombination creates new genome combinations by joining genome fragments of distinct “parental” origin. This phenomenon, frequent in viral...
  Recombination has an evident impact on virus evolution and emergence of new pathotypes, and has generated an immense literature. However, the distribution of...
SourceID plos
doaj
pubmedcentral
hal
proquest
gale
pubmed
crossref
SourceType Open Website
Open Access Repository
Aggregation Database
Index Database
Enrichment Source
StartPage e1002028
SubjectTerms Agrobacterium tumefaciens - genetics
Begomovirus
Begomovirus - classification
Begomovirus - genetics
Biology
Cloning
Cloning, Molecular
DNA, Viral - genetics
Escherichia coli - genetics
Experiments
Geminiviridae
Geminivirus
Gene Targeting
Genetic Association Studies
Genetic diversity
Genetic recombination
Genome, Viral
Genomes
Genomics
Life Sciences
Lycopersicon esculentum
Mutagenesis
Mutation
Phenotype
Phylogeny
Plasmids
Reassortant Viruses - classification
Reassortant Viruses - genetics
Recombination, Genetic
Sequence Analysis, DNA
Solanum lycopersicum - virology
Tomatoes
Viral genetics
Viruses
SummonAdditionalLinks – databaseName: Biological Science Database
  dbid: M7P
  link: http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwpV3db9MwELdYAYkXvscKA1kIiaewxE7t5AmVj2lIqKq2gfZm2a6zVlrj0LRD---5S9zSoA0eeKviS-OP8_nOPv9-hLzJwAVxea4jU-BuleAiMszZaJAhgUiexYUuGrIJORplZ2f5OGy41SGtcm0TG0M98Rb3yA_AjsLKBtEGe1_9iJA1Ck9XA4XGDrmNKAmsSd0bry0x2OeG-hSpcSLJhQhX57hMDsJIvasqvWxgSGPkY99amhoE_42d3plimmSvuvD1da7onxmVW0vU4YP_bdxDcj84p3TYatMjcsuVj8ndlq7y6gmxnxBlNxBkUV9QcB7peOpKv7yqZpa2SMg1lhzrcuLn9MjP8V2_qinGuXMIwxtNoCE9jJ7-9PT7bAHlJ5UDM1M_Jd8OP59-PIoCSUNkJRPLyMYmtYK5PLaJyViRGehJ-GG4MCzRWcGFniTGGud4nOXGDZiUsWXgeGlegLe2S3qlL90eoYOBNhwcTJ06mxrJjBSFyDX4iC6xNo37hK_HR9mAYI5EGheqOZaTEMm0_aRwVFUY1T6JNm9VLYLHP-Q_4NBvZBF_u3ngF-cqTGeFYRaof2YsOJTggeoUAnrrhM7cRHCT98lrVByFCBslpvCc61Vdqy8nIzVkSOQIcau8Uei4I_Q2CBUeGmt1uDYBXYbIXR3J_Y4k2Anb_doUm7rVrKPhV4XPIA5PYZ1JLpM-2UMdX3dMrX5rJvz9WnevL6abYvwy5u6VDvRLZUKC154O4ptFcuQxwFvaffKsnUibesK0ljxJoWayM8U6DemWlLNpg5POcQVl_PnfK_6C3GvPCTCJdZ_0louVe0nu2MvlrF68agzKL8tRfcI
  priority: 102
  providerName: ProQuest
– databaseName: Public Library of Science (PLoS) Journals Open Access
  dbid: FPL
  link: http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwlV3db9MwELdYAYkXvmGFgSyExFMgsVPbeSwfVZGmqhoD7c2yXWettCZR0w7tv-cuccsyrUK8Rblz4juf7d8l5ztC3iuAID7LTGRz_FoluIgs8y4aKCwgkqk4N3lTbEJOJursLJv-dRRv_MHnMvkUdPqxqsy6SRgKO-IBucu4EOhsjabH25UX1uNEhONx-1p2tp8mS_9uLT6YYyhkr7oo69vg5s2oyWvb0OjR_wrwmDwMgJMOWwt5Qu744im535agvHpG3FfMnBuKXtEypwAI6XTui3J9VS0cbbMb10g5McWsXNJxucS25aam6LsuwbVuRpeGkC96-rukvxYroDfF7X39nPwcfTv9Mo5C4YXISSbWkYtt6gTzWewSq1iuLHQZLiwXliVG5VyYWWKd9Z7HKrN-wKSMHQMwZXgOCOwF6RVl4Q8JHQyM5QAaTepdaiWzUuQiM4D7fOJcGvcJ346HdiErORbHuNDNrzYJ3kmrJ43q00F9fRLtWlVtVo5_8H_God7xYk7t5gaMkw5TVKPrBCatrAOQCKjSpOCkOy-M8jPBbdYn79BQNGbNKDAs59xs6lp__zHRQ4bFGcEXlXuZTjpMHwJTXoKwzoSjEKAyzMbV4TzqcMLcd923zVHUa2KNh8ca74FvncLekVwmfXKINr1VTK0BkgBIBMedweO3dn47me7I-GaMxys82JdWQgISTwfxfpYMaxPgyes-edlOnF0_AWZKnqTQM9mZUh1BupRiMW9yn3PcFRl_tV-m1-RB-90fg1KPSG-92vg35J67XC_q1dtmwfgDPJVqMA
  priority: 102
  providerName: Public Library of Science
Title Distribution of the Phenotypic Effects of Random Homologous Recombination between Two Virus Species
URI https://www.ncbi.nlm.nih.gov/pubmed/21573141
https://www.proquest.com/docview/1289101562
https://www.proquest.com/docview/867324450
https://www.proquest.com/docview/902362043
https://hal.inrae.fr/hal-02647631
https://pubmed.ncbi.nlm.nih.gov/PMC3088723
https://doaj.org/article/6758ebe8bc264610a4523ce6a8ed63b9
http://dx.doi.org/10.1371/journal.ppat.1002028
Volume 7
WOSCitedRecordID wos000291014000014&url=https%3A%2F%2Fcvtisr.summon.serialssolutions.com%2F%23%21%2Fsearch%3Fho%3Df%26include.ft.matches%3Dt%26l%3Dnull%26q%3D
hasFullText 1
inHoldings 1
isFullTextHit
isPrint
journalDatabaseRights – providerCode: PRVAON
  databaseName: DOAJ Directory of Open Access Journals
  customDbUrl:
  eissn: 1553-7374
  dateEnd: 99991231
  omitProxy: false
  ssIdentifier: ssj0041316
  issn: 1553-7374
  databaseCode: DOA
  dateStart: 20050101
  isFulltext: true
  titleUrlDefault: https://www.doaj.org/
  providerName: Directory of Open Access Journals
– providerCode: PRVPQU
  databaseName: Biological Science Database
  customDbUrl:
  eissn: 1553-7374
  dateEnd: 99991231
  omitProxy: false
  ssIdentifier: ssj0041316
  issn: 1553-7374
  databaseCode: M7P
  dateStart: 20050901
  isFulltext: true
  titleUrlDefault: http://search.proquest.com/biologicalscijournals
  providerName: ProQuest
– providerCode: PRVPQU
  databaseName: Health & Medical Collection
  customDbUrl:
  eissn: 1553-7374
  dateEnd: 99991231
  omitProxy: false
  ssIdentifier: ssj0041316
  issn: 1553-7374
  databaseCode: 7X7
  dateStart: 20050901
  isFulltext: true
  titleUrlDefault: https://search.proquest.com/healthcomplete
  providerName: ProQuest
– providerCode: PRVPQU
  databaseName: ProQuest Central
  customDbUrl:
  eissn: 1553-7374
  dateEnd: 99991231
  omitProxy: false
  ssIdentifier: ssj0041316
  issn: 1553-7374
  databaseCode: BENPR
  dateStart: 20050901
  isFulltext: true
  titleUrlDefault: https://www.proquest.com/central
  providerName: ProQuest
– providerCode: PRVPQU
  databaseName: Publicly Available Content Database
  customDbUrl:
  eissn: 1553-7374
  dateEnd: 99991231
  omitProxy: false
  ssIdentifier: ssj0041316
  issn: 1553-7374
  databaseCode: PIMPY
  dateStart: 20050901
  isFulltext: true
  titleUrlDefault: http://search.proquest.com/publiccontent
  providerName: ProQuest
– providerCode: PRVATS
  databaseName: Public Library of Science (PLoS) : Open Access Journals [open access]
  customDbUrl:
  eissn: 1553-7374
  dateEnd: 99991231
  omitProxy: false
  ssIdentifier: ssj0041316
  issn: 1553-7374
  databaseCode: FPL
  dateStart: 20050101
  isFulltext: true
  titleUrlDefault: http://www.plos.org/publications/
  providerName: Public Library of Science
link http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwrV3db9MwELdYAYkXxPcKo7IQEk9hSZzazmMHqzZpVNEAqTxZtufQSmtSNWnR_nvuErdq0Ka98BJVuUsd3118v0vOd4R8lABBXJrqwOT4toozHpjY2WAosYFIKsNc502zCTGZyOk0zfZafWFOWFseuBXcMQJaGEgaC64bfL1OIHSyjmvprjgzzda9UKTbYKpdg2FlbpqeYlOcQDDO_aY5JqJjr6PPy6WumwKkIXZi33NKTe3-3Qp9MMMEyd7yuqxuA6H_5lLuOafxM_LUo0o6amfznDxwxQvyuO0zefOS2K9YHtd3tqJlTgH1UUzuKuub5dxSn9SBFHBdV-WCzsoFXluuK4oh8wLi50aF1Od10fpPSTfzFdBxryaE26_Iz_Hpjy9nge-uEFgR8zqwoUksj10a2sjIOJcGBAE_DOMmjrTMGddXkbHGORbK1LhhLERoY0BMmuUAs16TXlEW7pDQ4VAbBshQJ84mRsRG8JynGsCdi6xNwj5hW_Eq60uPYweMa9V8TxMQgrRyUqgU5ZXSJ8HuqmVbeuMe_hPU3I4XC2c3J8CclDcndZ859ckH1LvC0hgF5t781uuqUuffJ2oUYwdGCDjFnUyXHaZPnikvYbJW-_0OIDIsudXhPOpwwgNuu6PNcKp70zobXSg8BwF0Ag4i2kR9cogmuhVMpQB3ABKE6DyGv9-a7e1kuiPjyJh0VziwLyW5ALidDMO7WVJsQIDbq_vkTfsc7O4TsKRgUQJ3JjpPSGciXUoxnzUFzhm6vpi9_R8KfUeetJ8BMEf1iPTq1dq9J4_spp5XqwE5EFPRHOWAPDw5nWSXg2YlgeM4uxhgKnAGlOz8W_brL50NeAA
linkProvider Directory of Open Access Journals
linkToHtml http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMw1V3Nb9MwFLe2DgQXvmGFARYCcQpL7DRODggNxtRqW1VtA42TsV1nrbQmoek29Z_ib-S9fJQGbXDagVsVvyR-7vt0nt-PkNchhCA2ipSjY9ytCnjgaGaN0wkRQCQK3VjFBdiE6PfD4-NosEJ-1mdhsKyytomFoR6mBvfIN8GOgmeDbIN9yH44iBqFX1drCI1SLHbt_AJStvx9bxv-3zeM7Xw--tR1KlQBxwgWzBzjat8EzEau8XTI4lCDi4UfmgeaeSqMeaCGnjbaWu6GkbYdJoRrGEQKiscQXsBzV8maj8LeImuD3v7gW237wSMUYKsIxuMIHgTVYT0uvM1KNt5lmZoVjU9dRIBfcoYFZsDCM6yOsDCzlZ2m-WXB7581nEtOcefu_7ac98idKvymW6W-3CcrNnlAbpaAnPOHxGxjH-EKAoymMYXwmA5GNkln82xsaNnrOceRA5UM0wntphO8Nz3LKWbyEz0ut1dpVQBHjy5S-nU8hfHDzIIhzR-RL9fC4WPSStLErhPa6SjNIYRWvjW-FkyLIA4iBVGw9Yzx3TbhtTxIU_VoR6iQU1l8eBSQq5XrJFGKZCVFbeIs7srKHiX_oP-IoragxQ7jxYV0eiIrgyUxkQQFD7WBkBlibOV3GDc2UKEdBlxHbfIKBVViD5EEi5RO1Fmey95hX24xhKqEzFxcSXTQIHpbEcUpMGtUdTAElgx7kzUoNxqUYAlN820jZHWJre7WnsRrLjABntQ799pkHXWqXphc_tYEeHytK5cP08UwvhmrExML8iXDQEBe4nfcq0kiRGrAc-ht8qRU3MU8IegW3PNhZqKh0g1GmiPJeFR0gucYIzD-9O8Tf0ludY_29-Rer7_7jNwuv4pgye4Gac2mZ_Y5uWHOZ-N8-qIyZ5R8v26V_wWgONwc
linkToPdf http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMw1V1Lb9QwELbaLSAuvKELBSIE4hSa2Fk7OSBUKFVXLatVW6pyMrbX6a7UJmGzbdW_xq9jJnGWBrVw6oHbKp487J3HN8l4PkJexwBBbJIoX6f4tooz7mtqjd-LkUAkiYNUpRXZhBgM4oODZLhAfjZ7YbCssvGJlaMe5Qbfka-CH4XIBtkGXU1dWcRwfeND8cNHBin80trQadQqsmXPzyB9K9_31-G_fkPpxue9T5u-YxjwjaB85ptAR4ZTmwQm1DFNYw3hFn5oxjUNVZwyrkahNtpaFsSJtj0qRGAooAbFUoAacN1FsgSQPKIdsjTsfxl-a-IARIeKeBWJeXzBOHcb95gIV52evCsKNauaoAbIBn8hMFb8AfMosTjGIs1OcZSXlwHhP-s5LwTIjbv_89LeI3ccLPfWaju6TxZs9oDcrIk6zx8Ss479hR01mJenHsBmbzi2WT47LybGq3tAlziyo7JRfuxt5sd4bn5SepjhH-tJ_drVc4Vx3t5Z7u1PpjC-W1hwsOUj8vVaZviYdLI8s8vE6_WUZgCtVWRNpAXVgqc8UYCObWhMFHQJa3RDGte7HSlEjmT1QVJADlevk0SNkk6jusSfn1XUvUv-If8R1W4ui53HqwP59FA6RyYxwQTDj7UBKA3YW0U9yozlKrYjznTSJa9QaSX2FslQlQ7VSVnK_u5ArlGksISMXVwptNMSeuuE0hwma5TbMAJLhj3LWpIrLUnwkKZ9tzFO9cK0Nte2JR4LYBIQYcPTsEuW0b6ahSnlb6uAyzd2c_mwNx_GO2PVYmZBv2TMBeQrUS-4WiRBBgfcn94lT2ojnj8ngHHBwgieTLTMuzWR9kg2GVcd4hliB8qe_v3BX5JbYOdyuz_YekZu1x9LsJJ3hXRm0xP7nNwwp7NJOX3hPJtHvl-3xf8CpmPk3A
openUrl ctx_ver=Z39.88-2004&ctx_enc=info%3Aofi%2Fenc%3AUTF-8&rfr_id=info%3Asid%2Fsummon.serialssolutions.com&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Distribution+of+the+phenotypic+effects+of+random+homologous+recombination+between+two+virus+species&rft.jtitle=PLoS+pathogens&rft.au=Florence+Vuillaume&rft.au=Ga%C3%ABl+Th%C3%A9baud&rft.au=Cica+Urbino&rft.au=Nad%C3%A8ge+Forfert&rft.date=2011-05-01&rft.pub=Public+Library+of+Science+%28PLoS%29&rft.issn=1553-7366&rft.eissn=1553-7374&rft.volume=7&rft.issue=5&rft.spage=e1002028&rft_id=info:doi/10.1371%2Fjournal.ppat.1002028&rft.externalDBID=DOA&rft.externalDocID=oai_doaj_org_article_6758ebe8bc264610a4523ce6a8ed63b9
thumbnail_l http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/lc.gif&issn=1553-7374&client=summon
thumbnail_m http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/mc.gif&issn=1553-7374&client=summon
thumbnail_s http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/sc.gif&issn=1553-7374&client=summon