Three types of rescue can avert extinction in a changing environment

Setting aside high-quality large areas of habitat to protect threatened populations is becoming increasingly difficult as humans fragment and degrade the environment. Biologists and managers therefore must determine the best way to shepherd small populations through the dual challenges of reductions...

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Vydáno v:Proceedings of the National Academy of Sciences - PNAS Ročník 112; číslo 33; s. 10557 - 10562
Hlavní autoři: Hufbauer, Ruth A, Szűcs, Marianna, Kasyon, Emily, Youngberg, Courtney, Koontz, Michael J, Richards, Christopher, Tuff, Ty, Melbourne, Brett A
Médium: Journal Article
Jazyk:angličtina
Vydáno: United States 18.08.2015
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ISSN:1091-6490
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Abstract Setting aside high-quality large areas of habitat to protect threatened populations is becoming increasingly difficult as humans fragment and degrade the environment. Biologists and managers therefore must determine the best way to shepherd small populations through the dual challenges of reductions in both the number of individuals and genetic variability. By bringing in additional individuals, threatened populations can be increased in size (demographic rescue) or provided with variation to facilitate adaptation and reduce inbreeding (genetic rescue). The relative strengths of demographic and genetic rescue for reducing extinction and increasing growth of threatened populations are untested, and which type of rescue is effective may vary with population size. Using the flour beetle (Tribolium castaneum) in a microcosm experiment, we disentangled the genetic and demographic components of rescue, and compared them with adaptation from standing genetic variation (evolutionary rescue in the strictest sense) using 244 experimental populations founded at either a smaller (50 individuals) or larger (150 individuals) size. Both types of rescue reduced extinction, and those effects were additive. Over the course of six generations, genetic rescue increased population sizes and intrinsic fitness substantially. Both large and small populations showed evidence of being able to adapt from standing genetic variation. Our results support the practice of genetic rescue in facilitating adaptation and reducing inbreeding depression, and suggest that demographic rescue alone may suffice in larger populations even if only moderately inbred individuals are available for addition.
AbstractList Setting aside high-quality large areas of habitat to protect threatened populations is becoming increasingly difficult as humans fragment and degrade the environment. Biologists and managers therefore must determine the best way to shepherd small populations through the dual challenges of reductions in both the number of individuals and genetic variability. By bringing in additional individuals, threatened populations can be increased in size (demographic rescue) or provided with variation to facilitate adaptation and reduce inbreeding (genetic rescue). The relative strengths of demographic and genetic rescue for reducing extinction and increasing growth of threatened populations are untested, and which type of rescue is effective may vary with population size. Using the flour beetle (Tribolium castaneum) in a microcosm experiment, we disentangled the genetic and demographic components of rescue, and compared them with adaptation from standing genetic variation (evolutionary rescue in the strictest sense) using 244 experimental populations founded at either a smaller (50 individuals) or larger (150 individuals) size. Both types of rescue reduced extinction, and those effects were additive. Over the course of six generations, genetic rescue increased population sizes and intrinsic fitness substantially. Both large and small populations showed evidence of being able to adapt from standing genetic variation. Our results support the practice of genetic rescue in facilitating adaptation and reducing inbreeding depression, and suggest that demographic rescue alone may suffice in larger populations even if only moderately inbred individuals are available for addition.
Author Kasyon, Emily
Tuff, Ty
Richards, Christopher
Hufbauer, Ruth A
Szűcs, Marianna
Youngberg, Courtney
Koontz, Michael J
Melbourne, Brett A
Author_xml – sequence: 1
  givenname: Ruth A
  surname: Hufbauer
  fullname: Hufbauer, Ruth A
  email: ruth.hufbauer@colostate.edu
  organization: Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, CO 80523-1177; Graduate Degree Program in Ecology, Colorado State University, Fort Collins, CO 80523; ruth.hufbauer@colostate.edu
– sequence: 2
  givenname: Marianna
  surname: Szűcs
  fullname: Szűcs, Marianna
  organization: Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, CO 80523-1177
– sequence: 3
  givenname: Emily
  surname: Kasyon
  fullname: Kasyon, Emily
  organization: Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, CO 80523-1177
– sequence: 4
  givenname: Courtney
  surname: Youngberg
  fullname: Youngberg, Courtney
  organization: Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, CO 80523-1177
– sequence: 5
  givenname: Michael J
  surname: Koontz
  fullname: Koontz, Michael J
  organization: Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, CO 80523-1177; Graduate Degree Program in Ecology, Colorado State University, Fort Collins, CO 80523; Graduate Group in Ecology, University of California, Davis, CA 95616
– sequence: 6
  givenname: Christopher
  surname: Richards
  fullname: Richards, Christopher
  organization: US Department of Agriculture-Agricultural Research Service National Center for Genetic Resources Preservation, Fort Collins, CO 80521
– sequence: 7
  givenname: Ty
  surname: Tuff
  fullname: Tuff, Ty
  organization: Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, CO 80309
– sequence: 8
  givenname: Brett A
  surname: Melbourne
  fullname: Melbourne, Brett A
  organization: Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, CO 80309
BackLink https://www.ncbi.nlm.nih.gov/pubmed/26240320$$D View this record in MEDLINE/PubMed
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Issue 33
Keywords migration
adaptation
extinction
genetic rescue
evolutionary rescue
Language English
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Snippet Setting aside high-quality large areas of habitat to protect threatened populations is becoming increasingly difficult as humans fragment and degrade the...
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StartPage 10557
SubjectTerms Animals
Biodiversity
Biological Evolution
Ecosystem
Environment
Evolution, Molecular
Extinction, Biological
Female
Genetic Fitness
Genetic Variation
Genetics, Population
Male
Microsatellite Repeats
Models, Genetic
Population Density
Probability
Tribolium - genetics
Title Three types of rescue can avert extinction in a changing environment
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