Activation‐induced deaminase–deficient MRL/lpr mice secrete high levels of protective antibodies against lupus nephritis

Objective We previously generated MRL/lpr mice deficient in activation‐induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice have high levels of unmutated (germline) autoreactive IgM, yet they experienced an increase in survival and an improvement in lupus...

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Vydané v:Arthritis & Rheumatism Ročník 63; číslo 4; s. 1086 - 1096
Hlavní autori: Jiang, Chuancang, Zhao, Ming‐Lang, Scearce, Richard M., Diaz, Marilyn
Médium: Journal Article
Jazyk:English
Vydavateľské údaje: Hoboken Wiley Subscription Services, Inc., A Wiley Company 01.04.2011
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ISSN:0004-3591, 1529-0131, 1529-0131
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Abstract Objective We previously generated MRL/lpr mice deficient in activation‐induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice have high levels of unmutated (germline) autoreactive IgM, yet they experienced an increase in survival and an improvement in lupus nephritis that exceeded that of MRL/lpr mice lacking IgG. The purpose of the present study was to test the hypothesis that high levels of germline autoreactive IgM in these mice confer protection against lupus nephritis. Methods Autoreactive IgM antibodies of various specificities, including antibodies against double‐stranded DNA (dsDNA), from AID‐deficient MRL/lpr mice were given to asymptomatic MRL/lpr mice, and the levels of cytokines, proteinuria, immune complex deposition in the kidneys, and glomerulonephritis were examined. Novel AID‐deficient MRL/lpr mice that lack any antibodies were generated for comparison to AID‐deficient MRL/lpr mice that secrete only IgM. Results Treatment with IgM anti‐dsDNA resulted in a dramatic improvement in lupus nephritis. Other autoreactive IgM antibodies, such as antiphospholipid and anti‐Sm, did not alter the pathologic changes. Secretion of proinflammatory cytokines by macrophages and the levels of inflammatory cells and apoptotic debris in the kidneys were lower in mice receiving IgM anti‐dsDNA. Protective IgM derived from AID‐deficient MRL/lpr mice displayed a distinct B cell repertoire, with a bias toward members of the VH7183 family. Conclusion IgM anti‐dsDNA protected MRL/lpr mice from lupus nephritis, likely by stopping the inflammatory cascade leading to kidney damage. A distinct repertoire of VH usage in IgM anti‐dsDNA hybridomas from AID‐deficient mice suggests that there is enrichment of a dedicated B cell population that secretes unmutated protective IgM in these mice.
AbstractList Objective We previously generated MRL/lpr mice deficient in activation-induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice have high levels of unmutated (germline) autoreactive IgM, yet they experienced an increase in survival and an improvement in lupus nephritis that exceeded that of MRL/lpr mice lacking IgG. The purpose of the present study was to test the hypothesis that high levels of germline autoreactive IgM in these mice confer protection against lupus nephritis. Methods Autoreactive IgM antibodies of various specificities, including antibodies against double-stranded DNA (dsDNA), from AID-deficient MRL/lpr mice were given to asymptomatic MRL/lpr mice, and the levels of cytokines, proteinuria, immune complex deposition in the kidneys, and glomerulonephritis were examined. Novel AID-deficient MRL/lpr mice that lack any antibodies were generated for comparison to AID-deficient MRL/lpr mice that secrete only IgM. Results Treatment with IgM anti-dsDNA resulted in a dramatic improvement in lupus nephritis. Other autoreactive IgM antibodies, such as antiphospholipid and anti-Sm, did not alter the pathologic changes. Secretion of proinflammatory cytokines by macrophages and the levels of inflammatory cells and apoptotic debris in the kidneys were lower in mice receiving IgM anti-dsDNA. Protective IgM derived from AID-deficient MRL/lpr mice displayed a distinct B cell repertoire, with a bias toward members of the VH7183 family. Conclusion IgM anti-dsDNA protected MRL/lpr mice from lupus nephritis, likely by stopping the inflammatory cascade leading to kidney damage. A distinct repertoire of VH usage in IgM anti-dsDNA hybridomas from AID-deficient mice suggests that there is enrichment of a dedicated B cell population that secretes unmutated protective IgM in these mice. [PUBLICATION ABSTRACT]
Objective We previously generated MRL/lpr mice deficient in activation‐induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice have high levels of unmutated (germline) autoreactive IgM, yet they experienced an increase in survival and an improvement in lupus nephritis that exceeded that of MRL/lpr mice lacking IgG. The purpose of the present study was to test the hypothesis that high levels of germline autoreactive IgM in these mice confer protection against lupus nephritis. Methods Autoreactive IgM antibodies of various specificities, including antibodies against double‐stranded DNA (dsDNA), from AID‐deficient MRL/lpr mice were given to asymptomatic MRL/lpr mice, and the levels of cytokines, proteinuria, immune complex deposition in the kidneys, and glomerulonephritis were examined. Novel AID‐deficient MRL/lpr mice that lack any antibodies were generated for comparison to AID‐deficient MRL/lpr mice that secrete only IgM. Results Treatment with IgM anti‐dsDNA resulted in a dramatic improvement in lupus nephritis. Other autoreactive IgM antibodies, such as antiphospholipid and anti‐Sm, did not alter the pathologic changes. Secretion of proinflammatory cytokines by macrophages and the levels of inflammatory cells and apoptotic debris in the kidneys were lower in mice receiving IgM anti‐dsDNA. Protective IgM derived from AID‐deficient MRL/lpr mice displayed a distinct B cell repertoire, with a bias toward members of the VH7183 family. Conclusion IgM anti‐dsDNA protected MRL/lpr mice from lupus nephritis, likely by stopping the inflammatory cascade leading to kidney damage. A distinct repertoire of VH usage in IgM anti‐dsDNA hybridomas from AID‐deficient mice suggests that there is enrichment of a dedicated B cell population that secretes unmutated protective IgM in these mice.
We previously generated MRL/lpr mice deficient in activation-induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice have high levels of unmutated (germline) autoreactive IgM, yet they experienced an increase in survival and an improvement in lupus nephritis that exceeded that of MRL/lpr mice lacking IgG. The purpose of the present study was to test the hypothesis that high levels of germline autoreactive IgM in these mice confer protection against lupus nephritis.OBJECTIVEWe previously generated MRL/lpr mice deficient in activation-induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice have high levels of unmutated (germline) autoreactive IgM, yet they experienced an increase in survival and an improvement in lupus nephritis that exceeded that of MRL/lpr mice lacking IgG. The purpose of the present study was to test the hypothesis that high levels of germline autoreactive IgM in these mice confer protection against lupus nephritis.Autoreactive IgM antibodies of various specificities, including antibodies against double-stranded DNA (dsDNA), from AID-deficient MRL/lpr mice were given to asymptomatic MRL/lpr mice, and the levels of cytokines, proteinuria, immune complex deposition in the kidneys, and glomerulonephritis were examined. Novel AID-deficient MRL/lpr mice that lack any antibodies were generated for comparison to AID-deficient MRL/lpr mice that secrete only IgM.METHODSAutoreactive IgM antibodies of various specificities, including antibodies against double-stranded DNA (dsDNA), from AID-deficient MRL/lpr mice were given to asymptomatic MRL/lpr mice, and the levels of cytokines, proteinuria, immune complex deposition in the kidneys, and glomerulonephritis were examined. Novel AID-deficient MRL/lpr mice that lack any antibodies were generated for comparison to AID-deficient MRL/lpr mice that secrete only IgM.Treatment with IgM anti-dsDNA resulted in a dramatic improvement in lupus nephritis. Other autoreactive IgM antibodies, such as antiphospholipid and anti-Sm, did not alter the pathologic changes. Secretion of proinflammatory cytokines by macrophages and the levels of inflammatory cells and apoptotic debris in the kidneys were lower in mice receiving IgM anti-dsDNA. Protective IgM derived from AID-deficient MRL/lpr mice displayed a distinct B cell repertoire, with a bias toward members of the V(H) 7183 family.RESULTSTreatment with IgM anti-dsDNA resulted in a dramatic improvement in lupus nephritis. Other autoreactive IgM antibodies, such as antiphospholipid and anti-Sm, did not alter the pathologic changes. Secretion of proinflammatory cytokines by macrophages and the levels of inflammatory cells and apoptotic debris in the kidneys were lower in mice receiving IgM anti-dsDNA. Protective IgM derived from AID-deficient MRL/lpr mice displayed a distinct B cell repertoire, with a bias toward members of the V(H) 7183 family.IgM anti-dsDNA protected MRL/lpr mice from lupus nephritis, likely by stopping the inflammatory cascade leading to kidney damage. A distinct repertoire of V(H) usage in IgM anti-dsDNA hybridomas from AID-deficient mice suggests that there is enrichment of a dedicated B cell population that secretes unmutated protective IgM in these mice.CONCLUSIONIgM anti-dsDNA protected MRL/lpr mice from lupus nephritis, likely by stopping the inflammatory cascade leading to kidney damage. A distinct repertoire of V(H) usage in IgM anti-dsDNA hybridomas from AID-deficient mice suggests that there is enrichment of a dedicated B cell population that secretes unmutated protective IgM in these mice.
We previously generated MRL/lpr mice deficient in activation-induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice have high levels of unmutated (germline) autoreactive IgM, yet they experienced an increase in survival and an improvement in lupus nephritis that exceeded that of MRL/lpr mice lacking IgG. The purpose of the present study was to test the hypothesis that high levels of germline autoreactive IgM in these mice confer protection against lupus nephritis. Autoreactive IgM antibodies of various specificities, including antibodies against double-stranded DNA (dsDNA), from AID-deficient MRL/lpr mice were given to asymptomatic MRL/lpr mice, and the levels of cytokines, proteinuria, immune complex deposition in the kidneys, and glomerulonephritis were examined. Novel AID-deficient MRL/lpr mice that lack any antibodies were generated for comparison to AID-deficient MRL/lpr mice that secrete only IgM. Treatment with IgM anti-dsDNA resulted in a dramatic improvement in lupus nephritis. Other autoreactive IgM antibodies, such as antiphospholipid and anti-Sm, did not alter the pathologic changes. Secretion of proinflammatory cytokines by macrophages and the levels of inflammatory cells and apoptotic debris in the kidneys were lower in mice receiving IgM anti-dsDNA. Protective IgM derived from AID-deficient MRL/lpr mice displayed a distinct B cell repertoire, with a bias toward members of the V(H) 7183 family. IgM anti-dsDNA protected MRL/lpr mice from lupus nephritis, likely by stopping the inflammatory cascade leading to kidney damage. A distinct repertoire of V(H) usage in IgM anti-dsDNA hybridomas from AID-deficient mice suggests that there is enrichment of a dedicated B cell population that secretes unmutated protective IgM in these mice.
Author Jiang, Chuancang
Zhao, Ming‐Lang
Diaz, Marilyn
Scearce, Richard M.
AuthorAffiliation 1 Laboratory of Molecular Genetics, D3-01, National Institute of Environmental Health Sciences, 111 TW Alexander Drive, Research Triangle Park, NC, 27709
3 Duke Human Vaccine Institute, 2 Genome Court, MSRBII 4102B, DUMC 103020, Durham, NC 27710
AuthorAffiliation_xml – name: 3 Duke Human Vaccine Institute, 2 Genome Court, MSRBII 4102B, DUMC 103020, Durham, NC 27710
– name: 1 Laboratory of Molecular Genetics, D3-01, National Institute of Environmental Health Sciences, 111 TW Alexander Drive, Research Triangle Park, NC, 27709
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  givenname: Ming‐Lang
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  givenname: Richard M.
  surname: Scearce
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  surname: Diaz
  fullname: Diaz, Marilyn
  email: diaz@niehs.nih.gov
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Cites_doi 10.1084/jem.189.11.1799
10.3109/08830189209055577
10.1073/pnas.0803644105
10.1016/S0065-2776(01)76021-X
10.1016/S0076-6879(83)03032-3
10.4049/jimmunol.178.8.4803
10.1002/eji.1830250146
10.1084/jem.20021560
10.1016/j.autrev.2009.02.035
10.1097/00002281-200309000-00005
10.1093/intimm/4.1.93
10.1016/j.autrev.2007.12.001
10.1172/JCI1817
10.4049/jimmunol.170.6.3223
10.1002/art.21379
10.1007/BF00351176
10.4049/jimmunol.178.11.7422
10.1146/annurev.immunol.19.1.275
10.1002/eji.1830220709
10.1016/S0092-8674(00)00078-7
10.3109/08916939009002973
10.4049/jimmunol.147.2.515
10.1084/jem.173.2.461
10.1517/14712590802681636
10.1084/jem.171.1.265
10.1016/S0065-2776(08)60342-9
10.1016/S1074-7613(00)80494-X
10.1191/0961203304lu485oa
10.1016/j.autrev.2008.07.047
10.4049/jimmunol.176.2.790
10.1016/j.clim.2003.10.007
10.4049/jimmunol.177.7.4727
10.1111/j.1749-6632.1995.tb55856.x
10.1016/j.imlet.2009.04.005
10.1084/jem.148.5.1198
10.1016/j.autrev.2007.11.024
10.1084/jem.20061446
10.1136/ard.58.2008.i49
10.1172/JCI119796
10.1080/08916930802629554
10.1084/jem.189.10.1639
10.4049/jimmunol.164.2.786
10.1111/j.1365-2249.2009.04058.x
10.1084/jem.20030132
10.4049/jimmunol.150.4.1591
10.1101/SQB.1989.054.01.108
10.1111/j.1365-2567.2008.02882.x
10.1073/pnas.97.3.1184
10.4049/jimmunol.165.8.4353
10.1097/00002281-200209000-00003
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Issue 4
Keywords Kidney disease
Immunopathology
Connective tissue disease
Skin disease
Urinary system disease
Antibody
Rodentia
Rheumatology
Autoimmune disease
Activation
Lupus nephritis
Prevention
Vertebrata
Mammalia
Mouse
Animal
Systemic disease
Language English
License http://onlinelibrary.wiley.com/termsAndConditions#vor
CC BY 4.0
Copyright © 2011 by the American College of Rheumatology.
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Notes Drs. Jiang and Zhao contributed equally to this work.
Drs. Jiang, Zhao, and Diaz have a pending patent application for the use of unmutated IgM antibodies in the treatment of lupus nephritis.
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Both of these authors contributed equally to this work
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PublicationCentury 2000
PublicationDate April 2011
PublicationDateYYYYMMDD 2011-04-01
PublicationDate_xml – month: 04
  year: 2011
  text: April 2011
PublicationDecade 2010
PublicationPlace Hoboken
PublicationPlace_xml – name: Hoboken
– name: Hoboken , NJ
– name: United States
– name: Atlanta
PublicationTitle Arthritis & Rheumatism
PublicationTitleAlternate Arthritis Rheum
PublicationYear 2011
Publisher Wiley Subscription Services, Inc., A Wiley Company
Wiley
Wiley Subscription Services, Inc
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– name: Wiley
– name: Wiley Subscription Services, Inc
References 2002; 14
1991; 173
2009; 42
2000; 5
2002; 196
2008; 7
2003; 15
2006; 176
2003; 170
2008; 105
1999; 189
2006; 177
2004; 199
2007; 178
2004; 110
1992; 8
1983; 103
1991; 147
1989; 54
2000; 102
1995; 25
2000; 76
2010; 159
1997; 100
1999; 58
2000; 97
2004; 13
1993; 150
2001; 19
1990; 171
2005; 52
2009; 9
1995; 764
2009; 8
2009; 124
2000; 164
2000; 165
1978; 148
1996; 5
2006; 203
1992; 22
1998; 101
1985; 37
2009; 126
1996; 23
1992; 4
e_1_2_7_5_2
e_1_2_7_3_2
e_1_2_7_9_2
e_1_2_7_7_2
e_1_2_7_19_2
e_1_2_7_17_2
e_1_2_7_15_2
e_1_2_7_13_2
e_1_2_7_41_2
e_1_2_7_11_2
e_1_2_7_43_2
e_1_2_7_45_2
e_1_2_7_47_2
e_1_2_7_26_2
e_1_2_7_49_2
e_1_2_7_28_2
e_1_2_7_50_2
e_1_2_7_25_2
e_1_2_7_23_2
e_1_2_7_31_2
e_1_2_7_21_2
e_1_2_7_33_2
e_1_2_7_35_2
e_1_2_7_37_2
e_1_2_7_39_2
e_1_2_7_4_2
e_1_2_7_2_2
e_1_2_7_8_2
e_1_2_7_6_2
e_1_2_7_18_2
e_1_2_7_16_2
e_1_2_7_14_2
e_1_2_7_40_2
e_1_2_7_12_2
e_1_2_7_42_2
e_1_2_7_10_2
e_1_2_7_44_2
e_1_2_7_46_2
e_1_2_7_48_2
e_1_2_7_27_2
Bloom DD (e_1_2_7_30_2) 1993; 150
e_1_2_7_29_2
Takahashi S (e_1_2_7_36_2) 1991; 147
e_1_2_7_24_2
e_1_2_7_51_2
e_1_2_7_22_2
e_1_2_7_32_2
e_1_2_7_20_2
e_1_2_7_34_2
e_1_2_7_38_2
References_xml – volume: 54
  start-page: 933
  year: 1989
  end-page: 46
  article-title: Structural patterns in anti‐DNA antibodies from MRL/lpr mice
  publication-title: Cold Spring Harb Symp Quant Biol
– volume: 105
  start-page: 9727
  year: 2008
  end-page: 32
  article-title: Autoreactive IgG memory antibodies in patients with systemic lupus erythematosus arise from nonreactive and polyreactive precursors
  publication-title: Proc Natl Acad Sci U S A
– volume: 5
  start-page: 181
  year: 1996
  end-page: 8
  article-title: Impaired IgG‐dependent anaphylaxis and Arthus reaction in FcγRIII (CD16) deficient mice
  publication-title: Immunity
– volume: 178
  start-page: 4803
  year: 2007
  end-page: 10
  article-title: Dendritic cells from lupus‐prone mice are defective in repressing immunoglobulin secretion
  publication-title: J Immunol
– volume: 177
  start-page: 4727
  year: 2006
  end-page: 34
  article-title: Activation of the lectin pathway by natural IgM in a model of ischemia/reperfusion injury
  publication-title: J Immunol
– volume: 148
  start-page: 1198
  year: 1978
  end-page: 215
  article-title: Spontaneous murine lupus‐like syndromes: clinical and immunopathological manifestations in several strains
  publication-title: J Exp Med
– volume: 124
  start-page: 70
  year: 2009
  end-page: 6
  article-title: Mouse monoclonal autoantibodies penetrate mouse macrophage cells and stimulate NF‐κB activation and TNF‐α release
  publication-title: Immunol Lett
– volume: 103
  start-page: 459
  year: 1983
  end-page: 69
  article-title: Production of monoclonal antibodies reacting with the cytoplasm and surface of differentiated cells
  publication-title: Methods Enzymol
– volume: 14
  start-page: 498
  year: 2002
  end-page: 503
  article-title: Mediators of injury in lupus nephritis
  publication-title: Curr Opin Rheumatol
– volume: 150
  start-page: 1591
  year: 1993
  end-page: 610
  article-title: V region gene analysis of anti‐Sm hybridomas from MRL/Mp‐lpr/lpr mice
  publication-title: J Immunol
– volume: 7
  start-page: 317
  year: 2008
  end-page: 21
  article-title: Anti‐DNA antibodies in the pathogenesis of lupus nephritis—the emerging mechanisms
  publication-title: Autoimmun Rev
– volume: 76
  start-page: 227
  year: 2000
  end-page: 324
  article-title: Systemic lupus erythematosus, complement deficiency, and apoptosis
  publication-title: Adv Immunol
– volume: 37
  start-page: 269
  year: 1985
  end-page: 390
  article-title: Murine models of systemic lupus erythematosus
  publication-title: Adv Immunol
– volume: 97
  start-page: 1184
  year: 2000
  end-page: 9
  article-title: Accelerated development of IgG autoantibodies and autoimmune disease in the absence of secreted IgM
  publication-title: Proc Natl Acad Sci U S A
– volume: 203
  start-page: 2255
  year: 2006
  end-page: 61
  article-title: Persistent expression of autoantibodies in SLE patients in remission
  publication-title: J Exp Med
– volume: 173
  start-page: 461
  year: 1991
  end-page: 70
  article-title: Somatic mutations in the variable regions of a human IgG anti‐double‐stranded DNA autoantibody suggest a role for antigen in the induction of systemic lupus erythematosus
  publication-title: J Exp Med
– volume: 8
  start-page: 209
  year: 2009
  end-page: 13
  article-title: Altered B cell receptor signaling in human systemic lupus erythematosus
  publication-title: Autoimmun Rev
– volume: 101
  start-page: 696
  year: 1998
  end-page: 702
  article-title: Loci predisposing to autoimmunity in MRL‐Fas lpr and C57BL/6‐Faslpr mice
  publication-title: J Clin Invest
– volume: 199
  start-page: 381
  year: 2004
  end-page: 98
  article-title: Pathogenic profiles and molecular signatures of antinuclear autoantibodies rescued from NZM2410 lupus mice
  publication-title: J Exp Med
– volume: 126
  start-page: 102
  year: 2009
  end-page: 13
  article-title: Activation‐induced deaminase heterozygous MRL/lpr mice are delayed in the production of high‐affinity pathogenic antibodies and in the development of lupus nephritis
  publication-title: Immunology
– volume: 42
  start-page: 89
  year: 2009
  end-page: 103
  article-title: Lupus‐prone MRL/fas‐lpr/lpr mice display increased AID expression and extensive DNA lesions, comprising deletions and insertions, in the immunoglobulin locus: concurrent upregulation of somatic hypermutation and class switch DNA recombination
  publication-title: Autoimmunity
– volume: 9
  start-page: 34
  year: 2009
  end-page: 9
  article-title: Interleukin 2 and systemic lupus erythematosus: beyond the transcriptional regulatory net abnormalities
  publication-title: Autoimmun Rev
– volume: 4
  start-page: 93
  year: 1992
  end-page: 9
  article-title: Selective pathogenicity of murine rheumatoid factors of the cryoprecipitable IgG3 subclass
  publication-title: Int Immunol
– volume: 164
  start-page: 786
  year: 2000
  end-page: 94
  article-title: Modulation of renal disease in MRL/lpr mice genetically deficient in the alternative complement pathway factor B
  publication-title: J Immunol
– volume: 58
  start-page: I49
  issue: Suppl 1
  year: 1999
  end-page: 55
  article-title: Tumour necrosis factor and other cytokines in murine lupus
  publication-title: Ann Rheum Dis
– volume: 147
  start-page: 515
  year: 1991
  end-page: 9
  article-title: IgG3 production in MRL/lpr mice is responsible for development of lupus nephritis
  publication-title: J Immunol
– volume: 110
  start-page: 22
  year: 2004
  end-page: 9
  article-title: Clinical, immunologic and genetic analysis of 29 patients with autosomal recessive hyper‐IgM syndrome due to activation‐induced cytidine deaminase deficiency
  publication-title: Clin Immunol
– volume: 7
  start-page: 309
  year: 2008
  end-page: 12
  article-title: Common variable immune deficiency (CVID) presenting as an autoimmune disease: role of memory B cells
  publication-title: Autoimmun Rev
– volume: 176
  start-page: 790
  year: 2006
  end-page: 802
  article-title: Early preplasma cells define a tolerance checkpoint for autoreactive B cells
  publication-title: J Immunol
– volume: 102
  start-page: 553
  year: 2000
  end-page: 63
  article-title: Class switch recombination and hypermutation require activation‐induced cytidine deaminase (AID), a potential RNA editing enzyme
  publication-title: Cell
– volume: 171
  start-page: 265
  year: 1990
  end-page: 92
  article-title: Anti‐DNA antibodies from autoimmune mice arise by clonal expansion and somatic mutation
  publication-title: J Exp Med
– volume: 764
  start-page: 410
  year: 1995
  end-page: 23
  article-title: B‐1 cellular origin and VH segment structure of IgG, IgA, and IgM anti‐DNA autoantibodies in patients with systemic lupus erythematosus
  publication-title: Ann N Y Acad Sci
– volume: 178
  start-page: 7422
  year: 2007
  end-page: 31
  article-title: Abrogation of lupus nephritis in activation‐induced deaminase‐deficient MRL/lpr mice
  publication-title: J Immunol
– volume: 100
  start-page: 2538
  year: 1997
  end-page: 46
  article-title: Lupus‐specific antibodies reveal an altered pattern of somatic mutation
  publication-title: J Clin Invest
– volume: 19
  start-page: 275
  year: 2001
  end-page: 90
  article-title: IgG Fc receptors
  publication-title: Annu Rev Immunol
– volume: 22
  start-page: 1719
  year: 1992
  end-page: 28
  article-title: Analysis of immunoglobulin variable region genes from human IgG anti‐DNA hybridomas
  publication-title: Eur J Immunol
– volume: 23
  start-page: 247
  year: 1996
  end-page: 51
  article-title: T cells in murine lupus: propagation and regulation of disease
  publication-title: Mol Biol Rep
– volume: 15
  start-page: 548
  year: 2003
  end-page: 56
  article-title: The central role of dendritic cells and interferon‐α in SLE
  publication-title: Curr Opin Rheumatol
– volume: 189
  start-page: 1639
  year: 1999
  end-page: 48
  article-title: A novel mouse with B cells but lacking serum antibody reveals an antibody‐independent role for B cells in murine lupus
  publication-title: J Exp Med
– volume: 52
  start-page: 3629
  year: 2005
  end-page: 38
  article-title: Inhibition of lupus disease by anti–double‐stranded DNA antibodies of the IgM isotype in the (NZB × NZW)F mouse
  publication-title: Arthritis Rheum
– volume: 196
  start-page: 1543
  year: 2002
  end-page: 52
  article-title: Anti‐DNA B cells in MRL/lpr mice show altered differentiation and editing pattern
  publication-title: J Exp Med
– volume: 170
  start-page: 3223
  year: 2003
  end-page: 32
  article-title: Lupus‐prone mice have an abnormal response to thioglycolate and an impaired clearance of apoptotic cells
  publication-title: J Immunol
– volume: 189
  start-page: 1799
  year: 1999
  end-page: 814
  article-title: MRL‐lpr/lpr mice exhibit a defect in maintaining developmental arrest and follicular exclusion of anti‐double‐stranded DNA B cells
  publication-title: J Exp Med
– volume: 25
  start-page: 279
  year: 1995
  end-page: 84
  article-title: The effect of VH residues 6 and 23 on IgG3 cryoprecipitation and glomerular deposition
  publication-title: Eur J Immunol
– volume: 8
  start-page: 247
  year: 1992
  end-page: 57
  article-title: Development and function of the early B cell repertoire
  publication-title: Int Rev Immunol
– volume: 5
  start-page: 147
  year: 2000
  end-page: 50
  article-title: Polyclonal B cell activation and antigen‐driven antibody response as mechanisms of autoantibody production in SLE
  publication-title: Autoimmunity
– volume: 165
  start-page: 4353
  year: 2000
  end-page: 8
  article-title: Unexpected autoantibody production in membrane Ig‐μ‐deficient/lpr mice
  publication-title: J Immunol
– volume: 13
  start-page: 36
  year: 2004
  end-page: 43
  article-title: Clinical significance of anti‐dsDNA antibody isotypes: IgG/IgM ratio of anti‐dsDNA antibodies as a prognostic marker for lupus nephritis
  publication-title: Lupus
– volume: 159
  start-page: 169
  year: 2010
  end-page: 75
  article-title: Deficiency in activation‐induced cytidine deaminase promotes systemic autoimmunity in lpr mice on a C57BL/6 background
  publication-title: Clin Exp Immunol
– volume: 9
  start-page: 201
  year: 2009
  end-page: 6
  article-title: Potential for anti‐DNA immunoglobulin peptide therapy in systemic lupus erythematosus
  publication-title: Expert Opin Biol Ther
– ident: e_1_2_7_10_2
  doi: 10.1084/jem.189.11.1799
– ident: e_1_2_7_50_2
  doi: 10.3109/08830189209055577
– ident: e_1_2_7_35_2
  doi: 10.1073/pnas.0803644105
– ident: e_1_2_7_12_2
  doi: 10.1016/S0065-2776(01)76021-X
– ident: e_1_2_7_28_2
  doi: 10.1016/S0076-6879(83)03032-3
– ident: e_1_2_7_18_2
  doi: 10.4049/jimmunol.178.8.4803
– ident: e_1_2_7_38_2
  doi: 10.1002/eji.1830250146
– ident: e_1_2_7_14_2
  doi: 10.1084/jem.20021560
– ident: e_1_2_7_19_2
  doi: 10.1016/j.autrev.2009.02.035
– ident: e_1_2_7_16_2
  doi: 10.1097/00002281-200309000-00005
– ident: e_1_2_7_37_2
  doi: 10.1093/intimm/4.1.93
– ident: e_1_2_7_8_2
  doi: 10.1016/j.autrev.2007.12.001
– ident: e_1_2_7_47_2
  doi: 10.1172/JCI1817
– ident: e_1_2_7_15_2
  doi: 10.4049/jimmunol.170.6.3223
– ident: e_1_2_7_49_2
  doi: 10.1002/art.21379
– ident: e_1_2_7_9_2
  doi: 10.1007/BF00351176
– ident: e_1_2_7_24_2
  doi: 10.4049/jimmunol.178.11.7422
– ident: e_1_2_7_40_2
  doi: 10.1146/annurev.immunol.19.1.275
– ident: e_1_2_7_3_2
  doi: 10.1002/eji.1830220709
– ident: e_1_2_7_25_2
  doi: 10.1016/S0092-8674(00)00078-7
– ident: e_1_2_7_20_2
  doi: 10.3109/08916939009002973
– volume: 147
  start-page: 515
  year: 1991
  ident: e_1_2_7_36_2
  article-title: IgG3 production in MRL/lpr mice is responsible for development of lupus nephritis
  publication-title: J Immunol
  doi: 10.4049/jimmunol.147.2.515
– ident: e_1_2_7_2_2
  doi: 10.1084/jem.173.2.461
– ident: e_1_2_7_32_2
  doi: 10.1517/14712590802681636
– ident: e_1_2_7_34_2
  doi: 10.1084/jem.171.1.265
– ident: e_1_2_7_23_2
  doi: 10.1016/S0065-2776(08)60342-9
– ident: e_1_2_7_39_2
  doi: 10.1016/S1074-7613(00)80494-X
– ident: e_1_2_7_42_2
  doi: 10.1191/0961203304lu485oa
– ident: e_1_2_7_6_2
  doi: 10.1016/j.autrev.2008.07.047
– ident: e_1_2_7_17_2
  doi: 10.4049/jimmunol.176.2.790
– ident: e_1_2_7_44_2
  doi: 10.1016/j.clim.2003.10.007
– ident: e_1_2_7_43_2
  doi: 10.4049/jimmunol.177.7.4727
– ident: e_1_2_7_4_2
  doi: 10.1111/j.1749-6632.1995.tb55856.x
– ident: e_1_2_7_48_2
  doi: 10.1016/j.imlet.2009.04.005
– ident: e_1_2_7_22_2
  doi: 10.1084/jem.148.5.1198
– ident: e_1_2_7_45_2
  doi: 10.1016/j.autrev.2007.11.024
– ident: e_1_2_7_41_2
  doi: 10.1084/jem.20061446
– ident: e_1_2_7_11_2
  doi: 10.1136/ard.58.2008.i49
– ident: e_1_2_7_5_2
  doi: 10.1172/JCI119796
– ident: e_1_2_7_31_2
  doi: 10.1080/08916930802629554
– ident: e_1_2_7_21_2
  doi: 10.1084/jem.189.10.1639
– ident: e_1_2_7_13_2
  doi: 10.4049/jimmunol.164.2.786
– ident: e_1_2_7_46_2
  doi: 10.1111/j.1365-2249.2009.04058.x
– ident: e_1_2_7_29_2
  doi: 10.1084/jem.20030132
– volume: 150
  start-page: 1591
  year: 1993
  ident: e_1_2_7_30_2
  article-title: V region gene analysis of anti‐Sm hybridomas from MRL/Mp‐lpr/lpr mice
  publication-title: J Immunol
  doi: 10.4049/jimmunol.150.4.1591
– ident: e_1_2_7_51_2
  doi: 10.1101/SQB.1989.054.01.108
– ident: e_1_2_7_26_2
  doi: 10.1111/j.1365-2567.2008.02882.x
– ident: e_1_2_7_27_2
  doi: 10.1073/pnas.97.3.1184
– ident: e_1_2_7_33_2
  doi: 10.4049/jimmunol.165.8.4353
– ident: e_1_2_7_7_2
  doi: 10.1097/00002281-200209000-00003
SSID ssj0002353
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Snippet Objective We previously generated MRL/lpr mice deficient in activation‐induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation....
We previously generated MRL/lpr mice deficient in activation-induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation. These mice...
Objective We previously generated MRL/lpr mice deficient in activation-induced deaminase (AID) that lack isotype switching and immunoglobulin hypermutation....
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StartPage 1086
SubjectTerms AICDA (Activation-Induced Cytidine Deaminase)
Animals
Antibodies - metabolism
Apoptosis - drug effects
Apoptosis - physiology
B-Lymphocytes - immunology
B-Lymphocytes - metabolism
Biological and medical sciences
Cytidine Deaminase - deficiency
Cytidine Deaminase - genetics
Cytidine Deaminase - metabolism
Disease Models, Animal
Diseases of the osteoarticular system
DNA - immunology
Hybridomas - immunology
Hybridomas - metabolism
Immunoglobulin M - metabolism
Immunoglobulin M - pharmacology
Kidney - immunology
Kidney - pathology
Kidneys
Lupus
Lupus Erythematosus, Systemic - complications
Lupus Erythematosus, Systemic - immunology
Lupus Erythematosus, Systemic - metabolism
Lupus Nephritis - metabolism
Lupus Nephritis - pathology
Lupus Nephritis - prevention & control
Medical research
Medical sciences
Mice
Mice, Inbred C57BL
Mice, Inbred MRL lpr
Mice, Knockout
Nephrology. Urinary tract diseases
Proteinuria - metabolism
Rodents
Urinary system involvement in other diseases. Miscellaneous
Title Activation‐induced deaminase–deficient MRL/lpr mice secrete high levels of protective antibodies against lupus nephritis
URI https://onlinelibrary.wiley.com/doi/abs/10.1002%2Fart.30230
https://www.ncbi.nlm.nih.gov/pubmed/21225690
https://www.proquest.com/docview/1517062948
https://www.proquest.com/docview/859757071
https://pubmed.ncbi.nlm.nih.gov/PMC3079514
Volume 63
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