Why elephant have trunks and giraffe long tongues: how plants shape large herbivore mouth morphology

We investigated whether mass and morphological spatial patterns in plants possibly induced the development of enlarged soft mouth parts in especially megaherbivores. We used power functions and geometric principles to explore allometric relationships of both morphological and foraging characteristic...

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Vydáno v:Acta zoologica (Stockholm) Ročník 97; číslo 2; s. 246 - 254
Hlavní autoři: Pretorius, Yolanda, de Boer, Willem F., Kortekaas, Kim, van Wijngaarden, Machiel, Grant, Rina C., Kohi, Edward M., Mwakiwa, Emmanuel, Slotow, Rob, Prins, Herbert H. T.
Médium: Journal Article
Jazyk:angličtina
Vydáno: Oxford Blackwell Publishing Ltd 01.04.2016
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ISSN:0001-7272, 1463-6395
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Abstract We investigated whether mass and morphological spatial patterns in plants possibly induced the development of enlarged soft mouth parts in especially megaherbivores. We used power functions and geometric principles to explore allometric relationships of both morphological and foraging characteristics of mammalian herbivores in the South African savannah, covering a body size range of more than three orders magnitude. Our results show that, although intradental mouth volume scaled to a power slightly less than one to body mass, actual bite volume, as measured in the field, scaled to body mass with a factor closer to 1.75. However, when including the volume added to intradental mouth volume by soft mouth parts, such as tongue and lips (or trunks in elephant), mouth volume scaled linearly with actual bite volume and in a similar fashion as actual bite volume to body size. Bite mass and bite leaf mass scaled linearly with body size. We conclude that these scaling relationships indicate that large herbivores use their enlarged soft mouth parts to not only increase bite volume and thereby bite mass, but also select soft plant parts and thereby increase the leaf mass fraction per bite.
AbstractList We investigated whether mass and morphological spatial patterns in plants possibly induced the development of enlarged soft mouth parts in especially megaherbivores. We used power functions and geometric principles to explore allometric relationships of both morphological and foraging characteristics of mammalian herbivores in the South African savannah, covering a body size range of more than three orders magnitude. Our results show that, although intradental mouth volume scaled to a power slightly less than one to body mass, actual bite volume, as measured in the field, scaled to body mass with a factor closer to 1.75. However, when including the volume added to intradental mouth volume by soft mouth parts, such as tongue and lips (or trunks in elephant), mouth volume scaled linearly with actual bite volume and in a similar fashion as actual bite volume to body size. Bite mass and bite leaf mass scaled linearly with body size. We conclude that these scaling relationships indicate that large herbivores use their enlarged soft mouth parts to not only increase bite volume and thereby bite mass, but also select soft plant parts and thereby increase the leaf mass fraction per bite.
Author van Wijngaarden, Machiel
Kohi, Edward M.
Kortekaas, Kim
Grant, Rina C.
Mwakiwa, Emmanuel
de Boer, Willem F.
Pretorius, Yolanda
Slotow, Rob
Prins, Herbert H. T.
Author_xml – sequence: 1
  givenname: Yolanda
  surname: Pretorius
  fullname: Pretorius, Yolanda
  email: yolanda4wildlife@gmail.com
  organization: Centre for Wildlife Management, University of Pretoria, Private bag X20, Hatfield, 0028, Pretoria, South Africa
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  givenname: Willem F.
  surname: de Boer
  fullname: de Boer, Willem F.
  organization: Resource Ecology Group, Wageningen University, Droevendaalsesteeg 3a, 6708 PB, Wageningen, The Netherlands
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  givenname: Kim
  surname: Kortekaas
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  organization: Resource Ecology Group, Wageningen University, Droevendaalsesteeg 3a, 6708 PB, Wageningen, The Netherlands
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  givenname: Machiel
  surname: van Wijngaarden
  fullname: van Wijngaarden, Machiel
  organization: Resource Ecology Group, Wageningen University, Droevendaalsesteeg 3a, 6708 PB, Wageningen, The Netherlands
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  surname: Grant
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  givenname: Edward M.
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  fullname: Kohi, Edward M.
  organization: Resource Ecology Group, Wageningen University, Droevendaalsesteeg 3a, 6708 PB, Wageningen, The Netherlands
– sequence: 7
  givenname: Emmanuel
  surname: Mwakiwa
  fullname: Mwakiwa, Emmanuel
  organization: Resource Ecology Group, Wageningen University, Droevendaalsesteeg 3a, 6708 PB, Wageningen, The Netherlands
– sequence: 8
  givenname: Rob
  surname: Slotow
  fullname: Slotow, Rob
  organization: Biological and Conservation Sciences, Westville Campus, University of KwaZulu-Natal, Private Bag X54001, 4000, Durban, South Africa
– sequence: 9
  givenname: Herbert H. T.
  surname: Prins
  fullname: Prins, Herbert H. T.
  organization: Resource Ecology Group, Wageningen University, Droevendaalsesteeg 3a, 6708 PB, Wageningen, The Netherlands
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Snippet We investigated whether mass and morphological spatial patterns in plants possibly induced the development of enlarged soft mouth parts in especially...
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SubjectTerms allometry
Animal behavior
bite mass
Body size
Elephants
feeding station
Herbivores
instantaneous intake
Leaves
Leerstoelgroep Resource Ecology
Morphology
Mouth
PE&RC
Resource Ecology
Title Why elephant have trunks and giraffe long tongues: how plants shape large herbivore mouth morphology
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