Understanding the regulation of chronic wounds by tissue inhibitors of matrix metalloproteinases through mathematical modelling

Understanding the biochemistry and pharmacodynamics of chronic wounds is of key importance, due to the millions of people in the UK affected and the significant cost to the NHS. Chronic wounds are characterised by elevated concentrations of matrix metalloproteinases (MMPs) that destroy the surroundi...

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Published in:Journal of theoretical biology Vol. 604; p. 112083
Main Authors: Dari, Sonia, O’dea, Reuben D., Fadai, Nabil T.
Format: Journal Article
Language:English
Published: England Elsevier Ltd 07.05.2025
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ISSN:0022-5193, 1095-8541, 1095-8541
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Abstract Understanding the biochemistry and pharmacodynamics of chronic wounds is of key importance, due to the millions of people in the UK affected and the significant cost to the NHS. Chronic wounds are characterised by elevated concentrations of matrix metalloproteinases (MMPs) that destroy the surrounding extracellular matrix (ECM). However, fibroblasts can produce tissue inhibitors of MMPs (TIMPs) in order to regulate wound healing. Therefore, the role of TIMPs in both acute and chronic wounds needs to be properly understood in order to develop therapeutic treatments. In this work, we propose a reaction-diffusion system of four partial differential equations that describe the interaction of the ECM, fibroblasts, MMPs, and TIMPs in a wound. We observe that, subject to parameter sets corresponding to both acute and chronic wound healing, this mathematical model gives rise to travelling wave solutions. Using bifurcation analysis, we demonstrate that excessive degradation of the ECM results in the emergence of chronic wounds, and the reversal of these chronic wounds is prohibited for lower TIMP production values. These results are replicated within a simplified model obtained via a parameter sensitivity analysis. This model is further extended to more realistic spatial domains where we demonstrate the effectiveness of a therapeutic hydrogel containing TIMPs as a treatment for chronic wounds.
AbstractList Understanding the biochemistry and pharmacodynamics of chronic wounds is of key importance, due to the millions of people in the UK affected and the significant cost to the NHS. Chronic wounds are characterised by elevated concentrations of matrix metalloproteinases (MMPs) that destroy the surrounding extracellular matrix (ECM). However, fibroblasts can produce tissue inhibitors of MMPs (TIMPs) in order to regulate wound healing. Therefore, the role of TIMPs in both acute and chronic wounds needs to be properly understood in order to develop therapeutic treatments. In this work, we propose a reaction-diffusion system of four partial differential equations that describe the interaction of the ECM, fibroblasts, MMPs, and TIMPs in a wound. We observe that, subject to parameter sets corresponding to both acute and chronic wound healing, this mathematical model gives rise to travelling wave solutions. Using bifurcation analysis, we demonstrate that excessive degradation of the ECM results in the emergence of chronic wounds, and the reversal of these chronic wounds is prohibited for lower TIMP production values. These results are replicated within a simplified model obtained via a parameter sensitivity analysis. This model is further extended to more realistic spatial domains where we demonstrate the effectiveness of a therapeutic hydrogel containing TIMPs as a treatment for chronic wounds.
Understanding the biochemistry and pharmacodynamics of chronic wounds is of key importance, due to the millions of people in the UK affected and the significant cost to the NHS. Chronic wounds are characterised by elevated concentrations of matrix metalloproteinases (MMPs) that destroy the surrounding extracellular matrix (ECM). However, fibroblasts can produce tissue inhibitors of MMPs (TIMPs) in order to regulate wound healing. Therefore, the role of TIMPs in both acute and chronic wounds needs to be properly understood in order to develop therapeutic treatments. In this work, we propose a reaction-diffusion system of four partial differential equations that describe the interaction of the ECM, fibroblasts, MMPs, and TIMPs in a wound. We observe that, subject to parameter sets corresponding to both acute and chronic wound healing, this mathematical model gives rise to travelling wave solutions. Using bifurcation analysis, we demonstrate that excessive degradation of the ECM results in the emergence of chronic wounds, and the reversal of these chronic wounds is prohibited for lower TIMP production values. These results are replicated within a simplified model obtained via a parameter sensitivity analysis. This model is further extended to more realistic spatial domains where we demonstrate the effectiveness of a therapeutic hydrogel containing TIMPs as a treatment for chronic wounds.Understanding the biochemistry and pharmacodynamics of chronic wounds is of key importance, due to the millions of people in the UK affected and the significant cost to the NHS. Chronic wounds are characterised by elevated concentrations of matrix metalloproteinases (MMPs) that destroy the surrounding extracellular matrix (ECM). However, fibroblasts can produce tissue inhibitors of MMPs (TIMPs) in order to regulate wound healing. Therefore, the role of TIMPs in both acute and chronic wounds needs to be properly understood in order to develop therapeutic treatments. In this work, we propose a reaction-diffusion system of four partial differential equations that describe the interaction of the ECM, fibroblasts, MMPs, and TIMPs in a wound. We observe that, subject to parameter sets corresponding to both acute and chronic wound healing, this mathematical model gives rise to travelling wave solutions. Using bifurcation analysis, we demonstrate that excessive degradation of the ECM results in the emergence of chronic wounds, and the reversal of these chronic wounds is prohibited for lower TIMP production values. These results are replicated within a simplified model obtained via a parameter sensitivity analysis. This model is further extended to more realistic spatial domains where we demonstrate the effectiveness of a therapeutic hydrogel containing TIMPs as a treatment for chronic wounds.
ArticleNumber 112083
Author Fadai, Nabil T.
Dari, Sonia
O’dea, Reuben D.
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Cites_doi 10.1073/pnas.0909115106
10.1016/j.compbiomed.2023.107342
10.1089/107632704323061834
10.3390/biom10081169
10.1098/rspb.1996.0217
10.1177/096368979200100505
10.1074/jbc.M611500200
10.1016/0025-5564(96)00044-2
10.1016/S0893-9659(04)90128-0
10.1016/0893-9659(94)90022-1
10.1007/s11538-005-9022-3
10.3109/10715762.2014.920087
10.1007/s11538-011-9712-y
10.3389/fsysb.2022.876075
10.1073/pnas.0711642105
10.1016/j.jtbi.2008.04.011
10.1089/wound.2019.1132
10.1016/j.bbamcr.2017.04.015
10.1111/j.1523-1755.2004.00900.x
10.1016/j.cpc.2009.09.018
10.1063/1.1680571
10.1098/rspb.1990.0061
10.3389/fsysb.2022.962790
10.1006/jtbi.1999.0971
10.1046/j.1524-475X.2001.00278.x
10.1007/s11538-010-9514-7
10.1016/j.jtbi.2015.07.023
10.1177/0022034509359125
10.1007/s00018-016-2268-0
10.1016/S0025-5564(99)00005-X
10.1016/S0025-5564(00)00008-0
10.1529/biophysj.105.077610
10.1371/journal.pone.0050028
10.1063/1.4826955
10.1089/wound.2012.0416
10.1111/j.1524-475X.2007.00271.x
10.1088/0022-3727/45/44/445201
10.1088/1478-3975/abf425
10.1152/ajplung.00015.2014
10.1111/exd.13909
10.1002/mame.200900052
10.1016/S0741-5214(03)01022-X
10.1098/rsif.2008.0536
10.1016/0009-2509(96)00299-0
10.3390/ijms17122085
10.1371/journal.pone.0024029
10.1016/j.clindermatol.2010.03.009
10.1007/BF02460102
10.1093/imammb/16.4.369
10.1371/journal.pone.0201855
10.1016/j.bbamcr.2010.01.003
10.1098/rsif.2020.0950
10.1038/s41598-020-75683-5
10.1089/wound.2015.0635
10.1378/chest.102.4.1085
10.1126/science.276.5309.75
10.1016/S0022-5193(05)80715-5
10.1242/jcs.023820
10.1093/imammb/17.4.379
10.1111/jcmm.14316
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References Maini, McElwain, Leavesley (bib0045) 2004; 17
Kaouri, Christodoulou, Chakraborty, Mendez, Skourides, Ruiz-Baier (bib0037) 2022; 2
Amar, Smith, Fields (bib0001) 2017; 1864
Otero (bib0059) 2020
Pettet, Byrne, McElwain, Norbury (bib0061) 1996; 136
Chaplain, Byrne (bib0006) 1996; 8
Stepanova, Byrne, Maini, Alarcon (bib0073) 2023; 16
Friedman, Siewe (bib0028) 2020; 82
Dallon, Sherratt, Maini, Ferguson (bib0018) 2000; 17
Kirschner, D., 2008. Uncertainty and sensitivity functions and implementation. (accessed: 22.07.2024).
Murphy, Hall, Maini, McCue, McElwain (bib0053) 2012; 74
Sheardown, Cheng (bib0069) 1996; 51
Dale, Sherrat, Maini (bib0014) 1997; 59
Guo, DiPietro (bib0033) 2010; 89
Waugh, Sherratt (bib0081) 2006; 68
Olsen, Maini, Sherratt, Dallon (bib0057) 1999; 158
Wang, Liu, Liu, Lu (bib0079) 2013; 20
Schugart, Friedman, Zhao, Sen (bib0068) 2008; 105
Batra, Robinson, Mehner, Hockla, Miller, Radisky, Radisky (bib0002) 2012; 7
Russo (bib0064) 2020
Vempati, Karagiannis, Popel (bib0077) 2007; 282
Pardo, Selman, Ramirez, Ramos, Montano, Stricklin, Raghu (bib0060) 1992; 102
Sherratt, Murray (bib0070) 1990; 241
Saltelli, Annoni, Azzini, Campolongo, Ratto, Tarantola (bib0067) 2010; 181
Yang, Zhu, Cui, Li, Jin (bib0085) 2009; 294
Chen, Qin, Liu, Zhong, Jing, Wu, Peng, Li, Peng (bib0007) 2023; 17
McDougall, Dallon, Sherratt, Maini (bib0048) 2006; 364
Frykberg, Banks (bib0029) 2015; 4
Gaffney, Maini, McCaig, Zhao, Forrester (bib0031) 1999; 16
Chua, Laurent, In Laurent, editors (bib0008) 2006
Hunt, Torres, Bachar-Wikstrom, Wikstrom (bib0036) 2024; 7
Nee, McMorrow, Campbell, Slattery, Ryan (bib0055) 2004; 66
Wong, Seet, Bascom, Isfort, Bard (bib0083) 2020; 10
Zanca, Flegg, Osborne (bib0086) 2022; 2
Fan, Saha, Hanjaya-Putra (bib0023) 2021; 9
Flegg, Menon, Byrne, McElwain (bib0025) 2020; 82
.
Krejner, Litwiniuk, Grzela (bib0040) 2016; 3
Zhao, Liang, Clarke, Jackson, Xue (bib0087) 2016; 17
Department of Health and Social Care, Care, S., 2021. Advanced wound care: develop new treatments in the UK. Accessed 11/11/2021
Deng, Fan, Xiao, Tian, Zheng, Li, He (bib0020) 2024; 9
Proteintech, Mmp1 polyclonal antibody.
Tavakoli, Klar (bib0075) 2021; 10
Menon, Flegg, McCue, Schugart, Dawson, McElwain (bib0050) 2012; 279
Orazov, Sakiyama, Graves (bib0058) 2012; 45
Cumming, McElwain, Upton (bib0013) 2009; 7
Pettet, Chaplain, McElwain, Byrne (bib0062) 1996; 263
Sherratt, Murray (bib0071) 1992; 1
Buranasin, Mizutani, Iwasaki, Mahasarakham, Kido, Takeda, Izumi (bib0005) 2018; 13
Martin (bib0047) 1997; 276
Wearing, Sherratt (bib0082) 2000; 165
Brew, Nagase (bib0004) 2010; 1803
Fadai, Sachak-Patwa, Byrne, Maini, Bafadhel, Nicolau (bib0022) 2021; 18
Tranquillo, Murray (bib0076) 1992; 158
Gill, Parks (bib0032) 2007; 40
Bianchi, Painter, Sherratt (bib0003) 2015; 383
Xue, Friedman, Sen (bib0084) 2009; 106
Strogatz (bib0074) 2015
Dale, Sherratt, Maini (bib0015) 1994; 7
Monfared, Ertl, Rothbauer (bib0052) 2020; 10
Flegg, Menon, Maini, McElwain (bib0026) 2015; 6
Dallon, Sherratt, Maini (bib0017) 2001; 9
Marino, Hogue, Ray, Kirschner (bib0046) 2008; 254
Collier, Legant, Marmer, Lubman, Saffarian, Wakatsuki, Elson, Goldberg (bib0010) 2011; 6
Dallon, Sherratt, Maini (bib0016) 1999; 1999
Harbin, Sohutskay, Vanderlaan, Fontaine, Mendenhall, Fisher, Voytik-Harbin, Tepole (bib0034) 2023; 165
Li, Moeen Rezakhanlou, Chavez-Munoz, Lai, Ghahary (bib0042) 2009; 332
Nyugen, Mobashery, Chang (bib0056) 2016
Liu, Xiong, Baca-Regen, Nagase, Baxter (bib0043) 2003; 38
Chuang, Degendorfer, Davies (bib0009) 2014; 48
Haugh (bib0035) 2006; 90
Siddiqui, Bernstein (bib0072) 2010; 28
Maini, McElwain, Leavesley (bib0044) 2004; 10
Murphy, Buenzli, Tambyah, Thompson, Hugo, Baker, Simpson (bib0054) 2021; 18
Wang, Guerrero-Juarez, Qui, Du, Chen, Figueroa, Plikus, Nie (bib0080) 2019; 28
Kobayashi, Kim, Liu, Sugiura, Kohyama, Fang, Wen, Abe, Wang, Atkinson, Shipley, Senior, Rennard (bib0039) 2014; 306
Mi, Rivière, Clermont, Steed, Vodovotz (bib0051) 2007; 15
Cukier, Fortuin, Shuler, Petschek, Schaibly (bib0012) 1973; 59
Menon, Flegg (bib0049) 2021; 10
Sabino, Keller (bib0065) 2015; 2
Wallace, Basehore, Zito (bib0078) 2021
Ziraldo, Mi, An, Vodovotz (bib0088) 2013; 2
Fu, Zheng, Chen, Wu, Yang, Chen, Song (bib0030) 2022; 13
Flegg, Byrne, McElwain (bib0024) 2010; 72
Dari, Fadai, O’dea (bib0019) 2023; 85
Sacco, Belviso, Romano, Carfora, Schonauer, Nurzynska, Montagnani, Meglio, Castaldo (bib0066) 2019; 23
Landen, Li, Stahle (bib0041) 2016; 73
Frantz, Stewart, Weaver (bib0027) 2010; 123
Cordiero, Bhattacharya, Schultz, Khaw (bib0011) 2000; 41
Batra (10.1016/j.jtbi.2025.112083_bib0002) 2012; 7
Dari (10.1016/j.jtbi.2025.112083_bib0019) 2023; 85
Fadai (10.1016/j.jtbi.2025.112083_bib0022) 2021; 18
10.1016/j.jtbi.2025.112083_bib0038
Marino (10.1016/j.jtbi.2025.112083_bib0046) 2008; 254
Nee (10.1016/j.jtbi.2025.112083_bib0055) 2004; 66
Flegg (10.1016/j.jtbi.2025.112083_bib0024) 2010; 72
Monfared (10.1016/j.jtbi.2025.112083_bib0052) 2020; 10
Zanca (10.1016/j.jtbi.2025.112083_bib0086) 2022; 2
Bianchi (10.1016/j.jtbi.2025.112083_bib0003) 2015; 383
Sherratt (10.1016/j.jtbi.2025.112083_bib0070) 1990; 241
Frantz (10.1016/j.jtbi.2025.112083_bib0027) 2010; 123
Russo (10.1016/j.jtbi.2025.112083_bib0064) 2020
Yang (10.1016/j.jtbi.2025.112083_bib0085) 2009; 294
Murphy (10.1016/j.jtbi.2025.112083_bib0054) 2021; 18
Brew (10.1016/j.jtbi.2025.112083_bib0004) 2010; 1803
Deng (10.1016/j.jtbi.2025.112083_bib0020) 2024; 9
Friedman (10.1016/j.jtbi.2025.112083_bib0028) 2020; 82
Wearing (10.1016/j.jtbi.2025.112083_bib0082) 2000; 165
Hunt (10.1016/j.jtbi.2025.112083_bib0036) 2024; 7
Chen (10.1016/j.jtbi.2025.112083_bib0007) 2023; 17
Kaouri (10.1016/j.jtbi.2025.112083_bib0037) 2022; 2
Flegg (10.1016/j.jtbi.2025.112083_bib0025) 2020; 82
Sacco (10.1016/j.jtbi.2025.112083_bib0066) 2019; 23
Wallace (10.1016/j.jtbi.2025.112083_bib0078) 2021
Cumming (10.1016/j.jtbi.2025.112083_bib0013) 2009; 7
Dallon (10.1016/j.jtbi.2025.112083_bib0018) 2000; 17
Otero (10.1016/j.jtbi.2025.112083_bib0059) 2020
Vempati (10.1016/j.jtbi.2025.112083_bib0077) 2007; 282
Dallon (10.1016/j.jtbi.2025.112083_bib0017) 2001; 9
Dale (10.1016/j.jtbi.2025.112083_bib0014) 1997; 59
Maini (10.1016/j.jtbi.2025.112083_bib0044) 2004; 10
Guo (10.1016/j.jtbi.2025.112083_bib0033) 2010; 89
Xue (10.1016/j.jtbi.2025.112083_bib0084) 2009; 106
Harbin (10.1016/j.jtbi.2025.112083_bib0034) 2023; 165
Liu (10.1016/j.jtbi.2025.112083_bib0043) 2003; 38
Pettet (10.1016/j.jtbi.2025.112083_bib0062) 1996; 263
Tavakoli (10.1016/j.jtbi.2025.112083_bib0075) 2021; 10
Buranasin (10.1016/j.jtbi.2025.112083_bib0005) 2018; 13
Cordiero (10.1016/j.jtbi.2025.112083_bib0011) 2000; 41
Wang (10.1016/j.jtbi.2025.112083_bib0080) 2019; 28
Chaplain (10.1016/j.jtbi.2025.112083_bib0006) 1996; 8
Sheardown (10.1016/j.jtbi.2025.112083_bib0069) 1996; 51
Gill (10.1016/j.jtbi.2025.112083_bib0032) 2007; 40
Pettet (10.1016/j.jtbi.2025.112083_bib0061) 1996; 136
Wang (10.1016/j.jtbi.2025.112083_bib0079) 2013; 20
Siddiqui (10.1016/j.jtbi.2025.112083_bib0072) 2010; 28
Ziraldo (10.1016/j.jtbi.2025.112083_bib0088) 2013; 2
Dale (10.1016/j.jtbi.2025.112083_bib0015) 1994; 7
Cukier (10.1016/j.jtbi.2025.112083_bib0012) 1973; 59
Wong (10.1016/j.jtbi.2025.112083_bib0083) 2020; 10
Zhao (10.1016/j.jtbi.2025.112083_bib0087) 2016; 17
Fan (10.1016/j.jtbi.2025.112083_bib0023) 2021; 9
Orazov (10.1016/j.jtbi.2025.112083_bib0058) 2012; 45
Schugart (10.1016/j.jtbi.2025.112083_bib0068) 2008; 105
Fu (10.1016/j.jtbi.2025.112083_bib0030) 2022; 13
Landen (10.1016/j.jtbi.2025.112083_bib0041) 2016; 73
Menon (10.1016/j.jtbi.2025.112083_bib0049) 2021; 10
Olsen (10.1016/j.jtbi.2025.112083_bib0057) 1999; 158
Frykberg (10.1016/j.jtbi.2025.112083_bib0029) 2015; 4
Haugh (10.1016/j.jtbi.2025.112083_bib0035) 2006; 90
McDougall (10.1016/j.jtbi.2025.112083_bib0048) 2006; 364
Stepanova (10.1016/j.jtbi.2025.112083_bib0073) 2023; 16
Strogatz (10.1016/j.jtbi.2025.112083_bib0074) 2015
Chua (10.1016/j.jtbi.2025.112083_bib0008) 2006
Dallon (10.1016/j.jtbi.2025.112083_bib0016) 1999; 1999
Gaffney (10.1016/j.jtbi.2025.112083_bib0031) 1999; 16
Murphy (10.1016/j.jtbi.2025.112083_bib0053) 2012; 74
Li (10.1016/j.jtbi.2025.112083_bib0042) 2009; 332
Tranquillo (10.1016/j.jtbi.2025.112083_bib0076) 1992; 158
Menon (10.1016/j.jtbi.2025.112083_bib0050) 2012; 279
Krejner (10.1016/j.jtbi.2025.112083_bib0040) 2016; 3
Collier (10.1016/j.jtbi.2025.112083_bib0010) 2011; 6
Kobayashi (10.1016/j.jtbi.2025.112083_bib0039) 2014; 306
10.1016/j.jtbi.2025.112083_bib0063
Sabino (10.1016/j.jtbi.2025.112083_bib0065) 2015; 2
10.1016/j.jtbi.2025.112083_bib0021
Amar (10.1016/j.jtbi.2025.112083_bib0001) 2017; 1864
Flegg (10.1016/j.jtbi.2025.112083_bib0026) 2015; 6
Sherratt (10.1016/j.jtbi.2025.112083_bib0071) 1992; 1
Waugh (10.1016/j.jtbi.2025.112083_bib0081) 2006; 68
Chuang (10.1016/j.jtbi.2025.112083_bib0009) 2014; 48
Martin (10.1016/j.jtbi.2025.112083_bib0047) 1997; 276
Mi (10.1016/j.jtbi.2025.112083_bib0051) 2007; 15
Maini (10.1016/j.jtbi.2025.112083_bib0045) 2004; 17
Nyugen (10.1016/j.jtbi.2025.112083_bib0056) 2016
Pardo (10.1016/j.jtbi.2025.112083_bib0060) 1992; 102
Saltelli (10.1016/j.jtbi.2025.112083_bib0067) 2010; 181
References_xml – volume: 158
  start-page: 145
  year: 1999
  end-page: 170
  ident: bib0057
  article-title: Mathematical modelling of anisotropy in fibrous connective tissue
  publication-title: Math. Biosci.
– volume: 45
  year: 2012
  ident: bib0058
  article-title: Wound healing modeling: investigating ambient gas plasma treatment efficacy
  publication-title: J. Phys. D
– volume: 2
  start-page: 1
  year: 2015
  end-page: 8
  ident: bib0065
  article-title: Matrix metalloproteinases in impaired wound healing
  publication-title: Metalloproteinases Med.
– year: 2020
  ident: bib0064
  publication-title: Bio-responsive hydrogels for an in vitro brain cancer cell model: self-controlled inhibition of matrix metalloproteinase activity
– volume: 7
  start-page: 11
  year: 1994
  end-page: 14
  ident: bib0015
  article-title: The speed of cornea1 epithelial wound healing
  publication-title: Appl. Math. Lett.
– volume: 10
  start-page: 475
  year: 2004
  end-page: 482
  ident: bib0044
  article-title: Travelling wave model to interpret a wound-healing cell migration assay for human peritoneal mesothelial cells
  publication-title: Tissue Eng.
– volume: 383
  start-page: 61
  year: 2015
  end-page: 86
  ident: bib0003
  article-title: A mathematical model for lymphangiogenesis in normal and diabetic wounds
  publication-title: J. Theor. Biol.
– volume: 17
  start-page: 379
  year: 2000
  end-page: 393
  ident: bib0018
  article-title: Biological implications of a discrete mathematical model for collagen deposition and alignment in dermal wound repair
  publication-title: IMA J. Math. Appl. Med. Biol.
– volume: 74
  start-page: 1143
  year: 2012
  end-page: 1170
  ident: bib0053
  article-title: A fibrocontractive mechanochemical model of dermal wound closure incorporating realistic growth factor kinetics
  publication-title: Bull. Math. Biol.
– volume: 10
  year: 2020
  ident: bib0083
  article-title: Red-COLA1: a human fibroblast reporter cell line for type i collagen transcription
  publication-title: Sci. Rep.
– volume: 59
  start-page: 1077
  year: 1997
  end-page: 1100
  ident: bib0014
  article-title: Role of fibroblast migration in collagen fiber formation during fetal and adult dermal wound healing
  publication-title: Bull. Math. Biol.
– volume: 28
  start-page: 519
  year: 2010
  end-page: 526
  ident: bib0072
  article-title: Chronic wound infection: facts and controversies
  publication-title: Clin. Dermatol.
– volume: 4
  start-page: 560
  year: 2015
  end-page: 582
  ident: bib0029
  article-title: Challenges in the treatment of chronic wounds
  publication-title: Adv. Wound Care New Rochelle
– volume: 10
  start-page: 328
  year: 2021
  ident: bib0049
  article-title: Mathematical modeling can advance wound healing research
  publication-title: Adv. Wound Care (New Rochelle)
– reference: Department of Health and Social Care, Care, S., 2021. Advanced wound care: develop new treatments in the UK. Accessed 11/11/2021
– volume: 364
  start-page: 1385
  year: 2006
  end-page: 1405
  ident: bib0048
  article-title: Fibroblast migration and collagen deposition during dermal wound healing: mathematical modelling and clinical implications
  publication-title: Philos. Trans. A Math. Phys. Eng. Sci.
– reference: Kirschner, D., 2008. Uncertainty and sensitivity functions and implementation. (accessed: 22.07.2024).
– year: 2020
  ident: bib0059
  publication-title: Enzyme-responsive hydrogels: development, characterisation and on-demand modulation of protease activity
– volume: 13
  start-page: 1
  year: 2022
  end-page: 10
  ident: bib0030
  article-title: Role of matrix metalloproteinases in diabetic foot ulcers: potential therapeutic targets
  publication-title: Front. Pharmacol.
– volume: 10
  year: 2020
  ident: bib0052
  article-title: An on-chip wound healing assay fabricated by xurography for evaluation of dermal fibroblast cell migration and wound closure
  publication-title: Sci. Rep.
– year: 2016
  ident: bib0056
  article-title: Roles of Matrix Metalloproteinases in Mutaneous Wound Healing
– volume: 282
  start-page: 37585
  year: 2007
  end-page: 37596
  ident: bib0077
  article-title: A biochemical model of matrix metalloproteinase 9 activation and inhibition
  publication-title: J. Biol. Chem.
– volume: 90
  start-page: 2297
  year: 2006
  end-page: 2308
  ident: bib0035
  article-title: Deterministic model of dermal wound invasion incorporating receptor-mediated signal transduction and spatial gradient sensing
  publication-title: Biophys. J.
– volume: 66
  start-page: 1376
  year: 2004
  end-page: 1386
  ident: bib0055
  article-title: Tnf-alpha and il-1beta-mediated regulation of mmp-9 and timp-1 in renal proximal tubular cells
  publication-title: Kidney Int.
– volume: 9
  year: 2024
  ident: bib0020
  article-title: Tgf-
  publication-title: Signal Trans. Target. Ther.
– volume: 165
  year: 2023
  ident: bib0034
  article-title: Computational mechanobiology model evaluating healing of postoperative cavities following breast-conserving surgery
  publication-title: Comput. Biol. Med.
– volume: 41
  start-page: 756
  year: 2000
  end-page: 763
  ident: bib0011
  article-title: Tgf-b1, -b2, and -b3 in vitro: biphasic effects on tenon’s fibroblast contraction, proliferation, and migration
  publication-title: Investig. Ophthalmol. Vis. Sci.
– volume: 1
  start-page: 365
  year: 1992
  end-page: 371
  ident: bib0071
  article-title: Epidermal wound healing: the clinical implications of a simple mathematical model
  publication-title: Cell Transplant.
– volume: 13
  year: 2018
  ident: bib0005
  article-title: High glucose-induced oxidative stress impairs proliferation and migration of human gingival fibroblasts
  publication-title: PLoS One
– volume: 254
  start-page: 178
  year: 2008
  end-page: 196
  ident: bib0046
  article-title: A methodology for performing global uncertainty and sensitivity analysis in systems biology
  publication-title: J. Theor. Biol.
– volume: 73
  start-page: 3861
  year: 2016
  end-page: 3885
  ident: bib0041
  article-title: Transition from inflammation to proliferation: a critical step during wound healing
  publication-title: Cell. Mol. Life Sci.
– volume: 20
  year: 2013
  ident: bib0079
  article-title: Mathematical model of gas plasma applied to chronic wounds
  publication-title: Phys. Plasmas
– volume: 9
  start-page: 278
  year: 2001
  end-page: 286
  ident: bib0017
  article-title: Modeling the effects of transforming growth factor-beta on extracellular matrix alignment in dermal wound repair
  publication-title: Wound Repair. Regen.
– volume: 82
  year: 2020
  ident: bib0025
  article-title: A current perspective on wound healing and tumour-induced angiogenesis
  publication-title: Bull. Math. Biol.
– volume: 123
  start-page: 4195
  year: 2010
  end-page: 4200
  ident: bib0027
  article-title: The extracellular matrix at a glance
  publication-title: J. Cell Sci.
– volume: 2
  start-page: 527
  year: 2013
  end-page: 537
  ident: bib0088
  article-title: Computational modeling of inflammation and wound healing
  publication-title: Adv. Wound Care (New Rochelle)
– volume: 28
  start-page: 493
  year: 2019
  end-page: 502
  ident: bib0080
  article-title: A multiscale hybrid mathematical model of epidermal-dermal interactions during skin wound healing
  publication-title: Exp. Dermatol.
– volume: 10
  start-page: 1169
  year: 2021
  end-page: 1189
  ident: bib0075
  article-title: Advanced hydrogels as wound dressings
  publication-title: Biomolecules
– volume: 7
  year: 2024
  ident: bib0036
  article-title: Cellular and molecular roles of reactive oxygen species in wound healing
  publication-title: Commun. Biol.
– volume: 15
  start-page: 672
  year: 2007
  end-page: 682
  ident: bib0051
  article-title: Agent-based model of inflammation and wound healing: insights into diabetic foot ulcer pathology and the role of transforming growth factor-
  publication-title: Wound Repair. Regen.
– volume: 1864
  start-page: 1940
  year: 2017
  end-page: 1951
  ident: bib0001
  article-title: Matrix metalloproteinase collagenolysis in health and disease
  publication-title: Biochim. Biophys. Acta Mol. Cell Res.
– volume: 16
  start-page: 369
  year: 1999
  end-page: 393
  ident: bib0031
  article-title: Modelling corneal epithelial wound closure in the presence of physiological electric fields via a moving boundary formalism
  publication-title: IMA J. Math. Appl. Med. Biol.
– year: 2015
  ident: bib0074
  article-title: Nonlinear Dynamics and Chaos
– volume: 102
  start-page: 1085
  year: 1992
  end-page: 1089
  ident: bib0060
  article-title: Production of collagenase and tissue inhibitor of metalloproteinases by fibroblasts derived from normal and fibrotic human lungs
  publication-title: Chest
– volume: 17
  year: 2016
  ident: bib0087
  article-title: Inflammation in chronic wounds
  publication-title: Int. J. Mol. Sci.
– volume: 9
  start-page: 1
  year: 2021
  end-page: 24
  ident: bib0023
  article-title: Biomimetic hydrogels to promote wound healing
  publication-title: Front. Bioeng. Biotechnol.
– year: 2021
  ident: bib0078
  article-title: Wound Healing Phases
– reference: Proteintech, Mmp1 polyclonal antibody.
– volume: 105
  start-page: 2628
  year: 2008
  end-page: 2633
  ident: bib0068
  article-title: Wound angiogenesis as a function of tissue oxygen tension: a mathematical model
  publication-title: Proc. Natl. Acad. Sci. USA
– volume: 85
  year: 2023
  ident: bib0019
  article-title: Modelling the effect of matrix metalloproteinases in dermal wound healing
  publication-title: Bull. Math. Biol.
– volume: 279
  start-page: 3329
  year: 2012
  end-page: 3338
  ident: bib0050
  article-title: Modelling the interaction of keratinocytes and fibroblasts during normal and abnormal wound healing processes
  publication-title: Proc. Biol. Sci.
– volume: 158
  start-page: 135
  year: 1992
  end-page: 172
  ident: bib0076
  article-title: Continuum model of fibroblast-driven wound contraction: inflammation-mediation
  publication-title: J. Theor. Biol.
– volume: 51
  start-page: 4517
  year: 1996
  end-page: 4529
  ident: bib0069
  article-title: Mechanisms of corneal epithelial wound healing
  publication-title: Chem. Eng. Sci.
– volume: 16
  year: 2023
  ident: bib0073
  article-title: Computational modeling of angiogenesis: the importance of cell rearrangements during vascular growth
  publication-title: WIREs Mech. Dis.
– volume: 165
  start-page: 41
  year: 2000
  end-page: 62
  ident: bib0082
  article-title: Keratinocyte growth factor signalling: a mathematical model of derma-epidermal interaction in epidermal wound healing
  publication-title: Math. Biosci.
– volume: 23
  start-page: 4256
  year: 2019
  end-page: 4268
  ident: bib0066
  article-title: Diversity of dermal fibroblasts as major determinant of variability in cell reprogramming
  publication-title: J. Cell Mol. Med.
– volume: 17
  year: 2023
  ident: bib0007
  article-title: Targeting matrix metalloproteases in diabetic wound healing
  publication-title: Front. Immunol.
– volume: 17
  start-page: 575
  year: 2004
  end-page: 580
  ident: bib0045
  article-title: Travelling waves in a wound healing assay
  publication-title: Appl. Math. Lett.
– volume: 7
  year: 2012
  ident: bib0002
  article-title: Pegylation extends circulation half-life while preserving in vitro and in vivo activity of tissue inhibitor of metalloproteinases-1 (timp-1)
  publication-title: PLoS One
– year: 2006
  ident: bib0008
  article-title: Encyclopedia of Respiratory Medicine Shapiro
  publication-title: Fibroblasts
– volume: 18
  start-page: 493
  year: 2021
  end-page: 502
  ident: bib0054
  article-title: The role of mechanical interactions in EMT
  publication-title: Phys. Biol.
– volume: 7
  start-page: 19
  year: 2009
  end-page: 34
  ident: bib0013
  article-title: A mathematical model of wound healing and subsequent scarring
  publication-title: J. R. Soc. Interface
– volume: 106
  start-page: 16782
  year: 2009
  end-page: 16787
  ident: bib0084
  article-title: A mathematical model of ischemic cutaneous wounds
  publication-title: PNAS
– volume: 59
  start-page: 3873
  year: 1973
  end-page: 3878
  ident: bib0012
  article-title: Study of sensitivity of coupled reaction systems to uncertainties in rate coefficients 1. theory
  publication-title: J. Chem. Phys.
– volume: 38
  start-page: 1376
  year: 2003
  end-page: 1383
  ident: bib0043
  article-title: Mechanism of inhibition of matrix metalloproteinase-2 expression by doxycycline in human aortic smooth muscle cells
  publication-title: J. Vasc. Surg.
– volume: 8
  start-page: 42
  year: 1996
  end-page: 48
  ident: bib0006
  article-title: Mathematical modelling of wound healing and tumour growth: two sides of the same coin
  publication-title: Wounds
– volume: 48
  start-page: 970
  year: 2014
  end-page: 989
  ident: bib0009
  article-title: Oxidation and modification of extracellular matrix and its role in disease
  publication-title: Free Radic. Res.
– volume: 72
  start-page: 1867
  year: 2010
  end-page: 1891
  ident: bib0024
  article-title: Mathematical model of hyperbaric oxygen therapy applied to chronic diabetic wounds
  publication-title: Bull. Math. Biol.
– volume: 276
  start-page: 75
  year: 1997
  end-page: 81
  ident: bib0047
  article-title: Wound healing - aiming for perfect skin regeneration
  publication-title: Science
– volume: 40
  start-page: 1334
  year: 2007
  end-page: 1347
  ident: bib0032
  article-title: Metalloproteinases and their inhibitors: regulators of wound healing
  publication-title: Int. J. Biochem. Cell Biol.
– volume: 136
  start-page: 35
  year: 1996
  end-page: 63
  ident: bib0061
  article-title: A model of wound-healing angiogenesis in soft tissue
  publication-title: Math. Biosci.
– volume: 3
  start-page: 29
  year: 2016
  end-page: 39
  ident: bib0040
  article-title: Matrix metalloproteinases in the wound microenvironment: therapeutic perspectives
  publication-title: Chronic Wound Care Manag. Res.
– volume: 1803
  start-page: 55
  year: 2010
  end-page: 71
  ident: bib0004
  article-title: The tissue inhibitors of metalloproteinases (TIMPs): an ancient family with structural and functional diversity
  publication-title: Biochim. Biophys. Acta
– volume: 241
  start-page: 29
  year: 1990
  end-page: 36
  ident: bib0070
  article-title: Models of epidermal wound healing
  publication-title: Proc. Biol. Sci.
– volume: 6
  start-page: 1
  year: 2011
  end-page: 14
  ident: bib0010
  article-title: Diffusion of MMPs on the surface of collagen fibrils: the mobile cell surface—collagen substratum interface
  publication-title: PLoS One
– volume: 82
  year: 2020
  ident: bib0028
  article-title: Mathematical model of chronic dermal wounds in diabetes and obesity
  publication-title: J. Theor. Biol.
– volume: 18
  year: 2021
  ident: bib0022
  article-title: Infection, inflammation and intervention: mechanistic modelling of epithelial cells in COVID-19
  publication-title: J. R. Soc. Interface
– volume: 89
  start-page: 219
  year: 2010
  end-page: 229
  ident: bib0033
  article-title: Factors affecting wound healing
  publication-title: J. Dent. Res.
– volume: 68
  start-page: 197
  year: 2006
  end-page: 207
  ident: bib0081
  article-title: Macrophage dynamics in diabetic wound dealing
  publication-title: Bull. Math. Biol.
– reference: .
– volume: 306
  start-page: 1006
  year: 2014
  end-page: 1015
  ident: bib0039
  article-title: Matrix metalloproteinase-9 activates TGF-
  publication-title: Am. J. Physiol. Lung Cell Mol. Physiol.
– volume: 2
  year: 2022
  ident: bib0037
  article-title: A new mechanochemical model for apical constriction: coupling calcium signalling and viscoelasticity
  publication-title: Front. Syst. Biol.
– volume: 294
  start-page: 611
  year: 2009
  end-page: 619
  ident: bib0085
  article-title: Electrospun composite mats of poly[(d,l-lactide)-co-glycolide] and collagen with high porosity as potential scaffolds for skin tissue engineering
  publication-title: Macromol. Mater. Eng.
– volume: 2
  year: 2022
  ident: bib0086
  article-title: Push or pull? cell proliferation and migration during wound healing
  publication-title: Front. Syst. Biol.
– volume: 1999
  start-page: 449
  year: 1999
  end-page: 471
  ident: bib0016
  article-title: Mathematical modelling of extracellular matrix dynamics using discrete cells: fiber orientation and tissue regeneration
  publication-title: J. Theor. Biol.
– volume: 332
  start-page: 1
  year: 2009
  end-page: 8
  ident: bib0042
  article-title: Keratinocyte-releasable factors increased the expression of MMP1 and MMP3 in co-cultured fibroblasts under both 2d and 3d culture conditions
  publication-title: Mol. Cell. Biochem.
– volume: 263
  start-page: 1487
  year: 1996
  end-page: 1493
  ident: bib0062
  article-title: On the role of angiogenesis in wound healing
  publication-title: Proc. R. Soc. Lond. B Biol. Sci.
– volume: 6
  start-page: 1
  year: 2015
  end-page: 17
  ident: bib0026
  article-title: On the mathematical modelling of wound healing angiogenesis in skin as a reaction-transport process
  publication-title: Front. Physiol.
– volume: 181
  start-page: 259
  year: 2010
  end-page: 270
  ident: bib0067
  article-title: Variance based sensitivity analysis of model output. design and estimator for the total sensitivity index
  publication-title: Comput. Phys. Commun.
– volume: 3
  start-page: 29
  year: 2016
  ident: 10.1016/j.jtbi.2025.112083_bib0040
  article-title: Matrix metalloproteinases in the wound microenvironment: therapeutic perspectives
  publication-title: Chronic Wound Care Manag. Res.
– volume: 8
  start-page: 42
  issue: 2
  year: 1996
  ident: 10.1016/j.jtbi.2025.112083_bib0006
  article-title: Mathematical modelling of wound healing and tumour growth: two sides of the same coin
  publication-title: Wounds
– volume: 40
  start-page: 1334
  issue: 6–7
  year: 2007
  ident: 10.1016/j.jtbi.2025.112083_bib0032
  article-title: Metalloproteinases and their inhibitors: regulators of wound healing
  publication-title: Int. J. Biochem. Cell Biol.
– volume: 106
  start-page: 16782
  issue: 39
  year: 2009
  ident: 10.1016/j.jtbi.2025.112083_bib0084
  article-title: A mathematical model of ischemic cutaneous wounds
  publication-title: PNAS
  doi: 10.1073/pnas.0909115106
– volume: 165
  year: 2023
  ident: 10.1016/j.jtbi.2025.112083_bib0034
  article-title: Computational mechanobiology model evaluating healing of postoperative cavities following breast-conserving surgery
  publication-title: Comput. Biol. Med.
  doi: 10.1016/j.compbiomed.2023.107342
– ident: 10.1016/j.jtbi.2025.112083_bib0021
– volume: 10
  start-page: 475
  issue: 3/4
  year: 2004
  ident: 10.1016/j.jtbi.2025.112083_bib0044
  article-title: Travelling wave model to interpret a wound-healing cell migration assay for human peritoneal mesothelial cells
  publication-title: Tissue Eng.
  doi: 10.1089/107632704323061834
– volume: 10
  start-page: 1169
  issue: 8
  year: 2021
  ident: 10.1016/j.jtbi.2025.112083_bib0075
  article-title: Advanced hydrogels as wound dressings
  publication-title: Biomolecules
  doi: 10.3390/biom10081169
– year: 2021
  ident: 10.1016/j.jtbi.2025.112083_bib0078
– volume: 9
  issue: 61
  year: 2024
  ident: 10.1016/j.jtbi.2025.112083_bib0020
  article-title: Tgf-β signaling in health, disease and therapeutics
  publication-title: Signal Trans. Target. Ther.
– volume: 2
  start-page: 1
  year: 2015
  ident: 10.1016/j.jtbi.2025.112083_bib0065
  article-title: Matrix metalloproteinases in impaired wound healing
  publication-title: Metalloproteinases Med.
– volume: 263
  start-page: 1487
  year: 1996
  ident: 10.1016/j.jtbi.2025.112083_bib0062
  article-title: On the role of angiogenesis in wound healing
  publication-title: Proc. R. Soc. Lond. B Biol. Sci.
  doi: 10.1098/rspb.1996.0217
– volume: 1
  start-page: 365
  issue: 5
  year: 1992
  ident: 10.1016/j.jtbi.2025.112083_bib0071
  article-title: Epidermal wound healing: the clinical implications of a simple mathematical model
  publication-title: Cell Transplant.
  doi: 10.1177/096368979200100505
– volume: 282
  start-page: 37585
  issue: 52
  year: 2007
  ident: 10.1016/j.jtbi.2025.112083_bib0077
  article-title: A biochemical model of matrix metalloproteinase 9 activation and inhibition
  publication-title: J. Biol. Chem.
  doi: 10.1074/jbc.M611500200
– volume: 82
  issue: 23
  year: 2020
  ident: 10.1016/j.jtbi.2025.112083_bib0025
  article-title: A current perspective on wound healing and tumour-induced angiogenesis
  publication-title: Bull. Math. Biol.
– volume: 136
  start-page: 35
  year: 1996
  ident: 10.1016/j.jtbi.2025.112083_bib0061
  article-title: A model of wound-healing angiogenesis in soft tissue
  publication-title: Math. Biosci.
  doi: 10.1016/0025-5564(96)00044-2
– volume: 17
  start-page: 575
  issue: 5
  year: 2004
  ident: 10.1016/j.jtbi.2025.112083_bib0045
  article-title: Travelling waves in a wound healing assay
  publication-title: Appl. Math. Lett.
  doi: 10.1016/S0893-9659(04)90128-0
– year: 2015
  ident: 10.1016/j.jtbi.2025.112083_bib0074
– volume: 279
  start-page: 3329
  issue: 1741
  year: 2012
  ident: 10.1016/j.jtbi.2025.112083_bib0050
  article-title: Modelling the interaction of keratinocytes and fibroblasts during normal and abnormal wound healing processes
  publication-title: Proc. Biol. Sci.
– volume: 7
  start-page: 11
  issue: 2
  year: 1994
  ident: 10.1016/j.jtbi.2025.112083_bib0015
  article-title: The speed of cornea1 epithelial wound healing
  publication-title: Appl. Math. Lett.
  doi: 10.1016/0893-9659(94)90022-1
– volume: 68
  start-page: 197
  issue: 1
  year: 2006
  ident: 10.1016/j.jtbi.2025.112083_bib0081
  article-title: Macrophage dynamics in diabetic wound dealing
  publication-title: Bull. Math. Biol.
  doi: 10.1007/s11538-005-9022-3
– volume: 48
  start-page: 970
  issue: 9
  year: 2014
  ident: 10.1016/j.jtbi.2025.112083_bib0009
  article-title: Oxidation and modification of extracellular matrix and its role in disease
  publication-title: Free Radic. Res.
  doi: 10.3109/10715762.2014.920087
– volume: 74
  start-page: 1143
  year: 2012
  ident: 10.1016/j.jtbi.2025.112083_bib0053
  article-title: A fibrocontractive mechanochemical model of dermal wound closure incorporating realistic growth factor kinetics
  publication-title: Bull. Math. Biol.
  doi: 10.1007/s11538-011-9712-y
– volume: 2
  year: 2022
  ident: 10.1016/j.jtbi.2025.112083_bib0086
  article-title: Push or pull? cell proliferation and migration during wound healing
  publication-title: Front. Syst. Biol.
  doi: 10.3389/fsysb.2022.876075
– volume: 105
  start-page: 2628
  issue: 7
  year: 2008
  ident: 10.1016/j.jtbi.2025.112083_bib0068
  article-title: Wound angiogenesis as a function of tissue oxygen tension: a mathematical model
  publication-title: Proc. Natl. Acad. Sci. USA
  doi: 10.1073/pnas.0711642105
– volume: 254
  start-page: 178
  issue: 1
  year: 2008
  ident: 10.1016/j.jtbi.2025.112083_bib0046
  article-title: A methodology for performing global uncertainty and sensitivity analysis in systems biology
  publication-title: J. Theor. Biol.
  doi: 10.1016/j.jtbi.2008.04.011
– ident: 10.1016/j.jtbi.2025.112083_bib0038
– volume: 10
  start-page: 328
  issue: 6
  year: 2021
  ident: 10.1016/j.jtbi.2025.112083_bib0049
  article-title: Mathematical modeling can advance wound healing research
  publication-title: Adv. Wound Care (New Rochelle)
  doi: 10.1089/wound.2019.1132
– year: 2020
  ident: 10.1016/j.jtbi.2025.112083_bib0064
– volume: 1864
  start-page: 1940
  issue: 11
  year: 2017
  ident: 10.1016/j.jtbi.2025.112083_bib0001
  article-title: Matrix metalloproteinase collagenolysis in health and disease
  publication-title: Biochim. Biophys. Acta Mol. Cell Res.
  doi: 10.1016/j.bbamcr.2017.04.015
– volume: 66
  start-page: 1376
  issue: 4
  year: 2004
  ident: 10.1016/j.jtbi.2025.112083_bib0055
  article-title: Tnf-alpha and il-1beta-mediated regulation of mmp-9 and timp-1 in renal proximal tubular cells
  publication-title: Kidney Int.
  doi: 10.1111/j.1523-1755.2004.00900.x
– volume: 181
  start-page: 259
  year: 2010
  ident: 10.1016/j.jtbi.2025.112083_bib0067
  article-title: Variance based sensitivity analysis of model output. design and estimator for the total sensitivity index
  publication-title: Comput. Phys. Commun.
  doi: 10.1016/j.cpc.2009.09.018
– volume: 59
  start-page: 3873
  issue: 8
  year: 1973
  ident: 10.1016/j.jtbi.2025.112083_bib0012
  article-title: Study of sensitivity of coupled reaction systems to uncertainties in rate coefficients 1. theory
  publication-title: J. Chem. Phys.
  doi: 10.1063/1.1680571
– volume: 241
  start-page: 29
  issue: 1300
  year: 1990
  ident: 10.1016/j.jtbi.2025.112083_bib0070
  article-title: Models of epidermal wound healing
  publication-title: Proc. Biol. Sci.
  doi: 10.1098/rspb.1990.0061
– volume: 2
  year: 2022
  ident: 10.1016/j.jtbi.2025.112083_bib0037
  article-title: A new mechanochemical model for apical constriction: coupling calcium signalling and viscoelasticity
  publication-title: Front. Syst. Biol.
  doi: 10.3389/fsysb.2022.962790
– volume: 1999
  start-page: 449
  issue: 4
  year: 1999
  ident: 10.1016/j.jtbi.2025.112083_bib0016
  article-title: Mathematical modelling of extracellular matrix dynamics using discrete cells: fiber orientation and tissue regeneration
  publication-title: J. Theor. Biol.
  doi: 10.1006/jtbi.1999.0971
– volume: 9
  start-page: 278
  issue: 4
  year: 2001
  ident: 10.1016/j.jtbi.2025.112083_bib0017
  article-title: Modeling the effects of transforming growth factor-beta on extracellular matrix alignment in dermal wound repair
  publication-title: Wound Repair. Regen.
  doi: 10.1046/j.1524-475X.2001.00278.x
– volume: 72
  start-page: 1867
  issue: 7
  year: 2010
  ident: 10.1016/j.jtbi.2025.112083_bib0024
  article-title: Mathematical model of hyperbaric oxygen therapy applied to chronic diabetic wounds
  publication-title: Bull. Math. Biol.
  doi: 10.1007/s11538-010-9514-7
– volume: 383
  start-page: 61
  year: 2015
  ident: 10.1016/j.jtbi.2025.112083_bib0003
  article-title: A mathematical model for lymphangiogenesis in normal and diabetic wounds
  publication-title: J. Theor. Biol.
  doi: 10.1016/j.jtbi.2015.07.023
– volume: 17
  issue: 14
  year: 2023
  ident: 10.1016/j.jtbi.2025.112083_bib0007
  article-title: Targeting matrix metalloproteases in diabetic wound healing
  publication-title: Front. Immunol.
– volume: 89
  start-page: 219
  issue: 3
  year: 2010
  ident: 10.1016/j.jtbi.2025.112083_bib0033
  article-title: Factors affecting wound healing
  publication-title: J. Dent. Res.
  doi: 10.1177/0022034509359125
– volume: 9
  start-page: 1
  issue: 718377
  year: 2021
  ident: 10.1016/j.jtbi.2025.112083_bib0023
  article-title: Biomimetic hydrogels to promote wound healing
  publication-title: Front. Bioeng. Biotechnol.
– volume: 16
  issue: 2
  year: 2023
  ident: 10.1016/j.jtbi.2025.112083_bib0073
  article-title: Computational modeling of angiogenesis: the importance of cell rearrangements during vascular growth
  publication-title: WIREs Mech. Dis.
– volume: 73
  start-page: 3861
  issue: 20
  year: 2016
  ident: 10.1016/j.jtbi.2025.112083_bib0041
  article-title: Transition from inflammation to proliferation: a critical step during wound healing
  publication-title: Cell. Mol. Life Sci.
  doi: 10.1007/s00018-016-2268-0
– volume: 158
  start-page: 145
  issue: 2
  year: 1999
  ident: 10.1016/j.jtbi.2025.112083_bib0057
  article-title: Mathematical modelling of anisotropy in fibrous connective tissue
  publication-title: Math. Biosci.
  doi: 10.1016/S0025-5564(99)00005-X
– volume: 165
  start-page: 41
  issue: 1
  year: 2000
  ident: 10.1016/j.jtbi.2025.112083_bib0082
  article-title: Keratinocyte growth factor signalling: a mathematical model of derma-epidermal interaction in epidermal wound healing
  publication-title: Math. Biosci.
  doi: 10.1016/S0025-5564(00)00008-0
– volume: 90
  start-page: 2297
  issue: 7
  year: 2006
  ident: 10.1016/j.jtbi.2025.112083_bib0035
  article-title: Deterministic model of dermal wound invasion incorporating receptor-mediated signal transduction and spatial gradient sensing
  publication-title: Biophys. J.
  doi: 10.1529/biophysj.105.077610
– volume: 85
  issue: 96
  year: 2023
  ident: 10.1016/j.jtbi.2025.112083_bib0019
  article-title: Modelling the effect of matrix metalloproteinases in dermal wound healing
  publication-title: Bull. Math. Biol.
– volume: 7
  issue: 11
  year: 2012
  ident: 10.1016/j.jtbi.2025.112083_bib0002
  article-title: Pegylation extends circulation half-life while preserving in vitro and in vivo activity of tissue inhibitor of metalloproteinases-1 (timp-1)
  publication-title: PLoS One
  doi: 10.1371/journal.pone.0050028
– volume: 364
  start-page: 1385
  issue: 1843
  year: 2006
  ident: 10.1016/j.jtbi.2025.112083_bib0048
  article-title: Fibroblast migration and collagen deposition during dermal wound healing: mathematical modelling and clinical implications
  publication-title: Philos. Trans. A Math. Phys. Eng. Sci.
– volume: 20
  year: 2013
  ident: 10.1016/j.jtbi.2025.112083_bib0079
  article-title: Mathematical model of gas plasma applied to chronic wounds
  publication-title: Phys. Plasmas
  doi: 10.1063/1.4826955
– volume: 2
  start-page: 527
  issue: 9
  year: 2013
  ident: 10.1016/j.jtbi.2025.112083_bib0088
  article-title: Computational modeling of inflammation and wound healing
  publication-title: Adv. Wound Care (New Rochelle)
  doi: 10.1089/wound.2012.0416
– volume: 82
  issue: 137
  year: 2020
  ident: 10.1016/j.jtbi.2025.112083_bib0028
  article-title: Mathematical model of chronic dermal wounds in diabetes and obesity
  publication-title: J. Theor. Biol.
– volume: 15
  start-page: 672
  issue: 5
  year: 2007
  ident: 10.1016/j.jtbi.2025.112083_bib0051
  article-title: Agent-based model of inflammation and wound healing: insights into diabetic foot ulcer pathology and the role of transforming growth factor-β1
  publication-title: Wound Repair. Regen.
  doi: 10.1111/j.1524-475X.2007.00271.x
– volume: 45
  issue: 44
  year: 2012
  ident: 10.1016/j.jtbi.2025.112083_bib0058
  article-title: Wound healing modeling: investigating ambient gas plasma treatment efficacy
  publication-title: J. Phys. D
  doi: 10.1088/0022-3727/45/44/445201
– volume: 41
  start-page: 756
  issue: 3
  year: 2000
  ident: 10.1016/j.jtbi.2025.112083_bib0011
  article-title: Tgf-b1, -b2, and -b3 in vitro: biphasic effects on tenon’s fibroblast contraction, proliferation, and migration
  publication-title: Investig. Ophthalmol. Vis. Sci.
– volume: 332
  start-page: 1
  issue: 1–2
  year: 2009
  ident: 10.1016/j.jtbi.2025.112083_bib0042
  article-title: Keratinocyte-releasable factors increased the expression of MMP1 and MMP3 in co-cultured fibroblasts under both 2d and 3d culture conditions
  publication-title: Mol. Cell. Biochem.
– volume: 18
  start-page: 493
  issue: 4
  year: 2021
  ident: 10.1016/j.jtbi.2025.112083_bib0054
  article-title: The role of mechanical interactions in EMT
  publication-title: Phys. Biol.
  doi: 10.1088/1478-3975/abf425
– volume: 306
  start-page: 1006
  issue: 11
  year: 2014
  ident: 10.1016/j.jtbi.2025.112083_bib0039
  article-title: Matrix metalloproteinase-9 activates TGF-β and stimulates fibroblast contraction of collagen gels
  publication-title: Am. J. Physiol. Lung Cell Mol. Physiol.
  doi: 10.1152/ajplung.00015.2014
– year: 2020
  ident: 10.1016/j.jtbi.2025.112083_bib0059
– volume: 28
  start-page: 493
  issue: 4
  year: 2019
  ident: 10.1016/j.jtbi.2025.112083_bib0080
  article-title: A multiscale hybrid mathematical model of epidermal-dermal interactions during skin wound healing
  publication-title: Exp. Dermatol.
  doi: 10.1111/exd.13909
– volume: 294
  start-page: 611
  issue: 9
  year: 2009
  ident: 10.1016/j.jtbi.2025.112083_bib0085
  article-title: Electrospun composite mats of poly[(d,l-lactide)-co-glycolide] and collagen with high porosity as potential scaffolds for skin tissue engineering
  publication-title: Macromol. Mater. Eng.
  doi: 10.1002/mame.200900052
– volume: 38
  start-page: 1376
  issue: 6
  year: 2003
  ident: 10.1016/j.jtbi.2025.112083_bib0043
  article-title: Mechanism of inhibition of matrix metalloproteinase-2 expression by doxycycline in human aortic smooth muscle cells
  publication-title: J. Vasc. Surg.
  doi: 10.1016/S0741-5214(03)01022-X
– volume: 7
  start-page: 19
  issue: 42
  year: 2009
  ident: 10.1016/j.jtbi.2025.112083_bib0013
  article-title: A mathematical model of wound healing and subsequent scarring
  publication-title: J. R. Soc. Interface
  doi: 10.1098/rsif.2008.0536
– volume: 51
  start-page: 4517
  issue: 19
  year: 1996
  ident: 10.1016/j.jtbi.2025.112083_bib0069
  article-title: Mechanisms of corneal epithelial wound healing
  publication-title: Chem. Eng. Sci.
  doi: 10.1016/0009-2509(96)00299-0
– volume: 10
  issue: 16192
  year: 2020
  ident: 10.1016/j.jtbi.2025.112083_bib0052
  article-title: An on-chip wound healing assay fabricated by xurography for evaluation of dermal fibroblast cell migration and wound closure
  publication-title: Sci. Rep.
– volume: 17
  issue: 12
  year: 2016
  ident: 10.1016/j.jtbi.2025.112083_bib0087
  article-title: Inflammation in chronic wounds
  publication-title: Int. J. Mol. Sci.
  doi: 10.3390/ijms17122085
– volume: 13
  start-page: 1
  issue: 1050630
  year: 2022
  ident: 10.1016/j.jtbi.2025.112083_bib0030
  article-title: Role of matrix metalloproteinases in diabetic foot ulcers: potential therapeutic targets
  publication-title: Front. Pharmacol.
– volume: 6
  start-page: 1
  issue: 9
  year: 2011
  ident: 10.1016/j.jtbi.2025.112083_bib0010
  article-title: Diffusion of MMPs on the surface of collagen fibrils: the mobile cell surface—collagen substratum interface
  publication-title: PLoS One
  doi: 10.1371/journal.pone.0024029
– volume: 28
  start-page: 519
  issue: 5
  year: 2010
  ident: 10.1016/j.jtbi.2025.112083_bib0072
  article-title: Chronic wound infection: facts and controversies
  publication-title: Clin. Dermatol.
  doi: 10.1016/j.clindermatol.2010.03.009
– volume: 59
  start-page: 1077
  issue: 6
  year: 1997
  ident: 10.1016/j.jtbi.2025.112083_bib0014
  article-title: Role of fibroblast migration in collagen fiber formation during fetal and adult dermal wound healing
  publication-title: Bull. Math. Biol.
  doi: 10.1007/BF02460102
– volume: 16
  start-page: 369
  issue: 4
  year: 1999
  ident: 10.1016/j.jtbi.2025.112083_bib0031
  article-title: Modelling corneal epithelial wound closure in the presence of physiological electric fields via a moving boundary formalism
  publication-title: IMA J. Math. Appl. Med. Biol.
  doi: 10.1093/imammb/16.4.369
– volume: 13
  issue: 8
  year: 2018
  ident: 10.1016/j.jtbi.2025.112083_bib0005
  article-title: High glucose-induced oxidative stress impairs proliferation and migration of human gingival fibroblasts
  publication-title: PLoS One
  doi: 10.1371/journal.pone.0201855
– year: 2006
  ident: 10.1016/j.jtbi.2025.112083_bib0008
  article-title: Encyclopedia of Respiratory Medicine Shapiro
– volume: 1803
  start-page: 55
  issue: 1
  year: 2010
  ident: 10.1016/j.jtbi.2025.112083_bib0004
  article-title: The tissue inhibitors of metalloproteinases (TIMPs): an ancient family with structural and functional diversity
  publication-title: Biochim. Biophys. Acta
  doi: 10.1016/j.bbamcr.2010.01.003
– volume: 18
  issue: 175
  year: 2021
  ident: 10.1016/j.jtbi.2025.112083_bib0022
  article-title: Infection, inflammation and intervention: mechanistic modelling of epithelial cells in COVID-19
  publication-title: J. R. Soc. Interface
  doi: 10.1098/rsif.2020.0950
– ident: 10.1016/j.jtbi.2025.112083_bib0063
– volume: 10
  issue: 1
  year: 2020
  ident: 10.1016/j.jtbi.2025.112083_bib0083
  article-title: Red-COLA1: a human fibroblast reporter cell line for type i collagen transcription
  publication-title: Sci. Rep.
  doi: 10.1038/s41598-020-75683-5
– volume: 6
  start-page: 1
  issue: 262
  year: 2015
  ident: 10.1016/j.jtbi.2025.112083_bib0026
  article-title: On the mathematical modelling of wound healing angiogenesis in skin as a reaction-transport process
  publication-title: Front. Physiol.
– volume: 4
  start-page: 560
  issue: 9
  year: 2015
  ident: 10.1016/j.jtbi.2025.112083_bib0029
  article-title: Challenges in the treatment of chronic wounds
  publication-title: Adv. Wound Care New Rochelle
  doi: 10.1089/wound.2015.0635
– volume: 7
  issue: 1534
  year: 2024
  ident: 10.1016/j.jtbi.2025.112083_bib0036
  article-title: Cellular and molecular roles of reactive oxygen species in wound healing
  publication-title: Commun. Biol.
– year: 2016
  ident: 10.1016/j.jtbi.2025.112083_bib0056
– volume: 102
  start-page: 1085
  issue: 4
  year: 1992
  ident: 10.1016/j.jtbi.2025.112083_bib0060
  article-title: Production of collagenase and tissue inhibitor of metalloproteinases by fibroblasts derived from normal and fibrotic human lungs
  publication-title: Chest
  doi: 10.1378/chest.102.4.1085
– volume: 276
  start-page: 75
  issue: 5309
  year: 1997
  ident: 10.1016/j.jtbi.2025.112083_bib0047
  article-title: Wound healing - aiming for perfect skin regeneration
  publication-title: Science
  doi: 10.1126/science.276.5309.75
– volume: 158
  start-page: 135
  issue: 2
  year: 1992
  ident: 10.1016/j.jtbi.2025.112083_bib0076
  article-title: Continuum model of fibroblast-driven wound contraction: inflammation-mediation
  publication-title: J. Theor. Biol.
  doi: 10.1016/S0022-5193(05)80715-5
– volume: 123
  start-page: 4195
  issue: 24
  year: 2010
  ident: 10.1016/j.jtbi.2025.112083_bib0027
  article-title: The extracellular matrix at a glance
  publication-title: J. Cell Sci.
  doi: 10.1242/jcs.023820
– volume: 17
  start-page: 379
  issue: 4
  year: 2000
  ident: 10.1016/j.jtbi.2025.112083_bib0018
  article-title: Biological implications of a discrete mathematical model for collagen deposition and alignment in dermal wound repair
  publication-title: IMA J. Math. Appl. Med. Biol.
  doi: 10.1093/imammb/17.4.379
– volume: 23
  start-page: 4256
  issue: 6
  year: 2019
  ident: 10.1016/j.jtbi.2025.112083_bib0066
  article-title: Diversity of dermal fibroblasts as major determinant of variability in cell reprogramming
  publication-title: J. Cell Mol. Med.
  doi: 10.1111/jcmm.14316
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Snippet Understanding the biochemistry and pharmacodynamics of chronic wounds is of key importance, due to the millions of people in the UK affected and the...
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SubjectTerms Chronic Disease
Extracellular Matrix - metabolism
Fibroblasts - metabolism
Humans
Matrix Metalloproteinases - metabolism
Models, Biological
Tissue Inhibitor of Metalloproteinases - metabolism
Wound Healing - physiology
Title Understanding the regulation of chronic wounds by tissue inhibitors of matrix metalloproteinases through mathematical modelling
URI https://dx.doi.org/10.1016/j.jtbi.2025.112083
https://www.ncbi.nlm.nih.gov/pubmed/40020775
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