Integrin αvβ6 critically regulates hepatic progenitor cell function and promotes ductular reaction, fibrosis, and tumorigenesis

Integrin αvβ6 is rapidly up‐regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent transforming growth factor beta 1 (TGFβ1). In human liver cirrhosis, αvβ6 is overexpressed by cells comprising the ductular reaction, and its inhibiti...

Celý popis

Uloženo v:
Podrobná bibliografie
Vydáno v:Hepatology (Baltimore, Md.) Ročník 63; číslo 1; s. 217 - 232
Hlavní autoři: Peng, Zhen‐Wei, Ikenaga, Naoki, Liu, Susan B., Sverdlov, Deanna Y., Vaid, Kahini A., Dixit, Richa, Weinreb, Paul H., Violette, Shelia, Sheppard, Dean, Schuppan, Detlef, Popov, Yury
Médium: Journal Article
Jazyk:angličtina
Vydáno: United States 01.01.2016
Témata:
ISSN:0270-9139, 1527-3350
On-line přístup:Získat plný text
Tagy: Přidat tag
Žádné tagy, Buďte první, kdo vytvoří štítek k tomuto záznamu!
Abstract Integrin αvβ6 is rapidly up‐regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent transforming growth factor beta 1 (TGFβ1). In human liver cirrhosis, αvβ6 is overexpressed by cells comprising the ductular reaction, and its inhibition suppresses experimental biliary fibrosis in rodents. Here, we show that αvβ6 is expressed on the actively proliferating subset of hepatic progenitor cells and is required for their progenitor function in vivo and in vitro through integrin αvβ6‐dependent TGFβ1 activation. Freshly isolated αvβ6+ liver cells demonstrate clonogenic potential and differentiate into cholangiocytes and functional hepatocytes in vitro, whereas colony formation by epithelial cell adhesion molecule‐positive progenitor cells is blocked by αvβ6‐neutralizing antibody and in integrin beta 6‐deficient cells. Inhibition of progenitors by anti‐αvβ6 antibody is recapitulated by TGFβ1 neutralization and rescued by addition of bioactive TGFβ1. Genetic disruption or selective targeting of αvβ6 with 3G9 antibody potently inhibits progenitor cell responses in mouse models of chronic biliary injury and protects from liver fibrosis and tumorigenesis, two conditions clinically associated with exacerbated ductular reaction. Conclusion: These results suggest that αvβ6 is a promising target for chronic fibrotic liver diseases and associated cancers. (Hepatology 2016;63:217–232)
AbstractList Integrin αvβ6 is rapidly up-regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent transforming growth factor beta 1 (TGFβ1). In human liver cirrhosis, αvβ6 is overexpressed by cells comprising the ductular reaction, and its inhibition suppresses experimental biliary fibrosis in rodents. Here, we show that αvβ6 is expressed on the actively proliferating subset of hepatic progenitor cells and is required for their progenitor function in vivo and in vitro through integrin αvβ6-dependent TGFβ1 activation. Freshly isolated αvβ6(+) liver cells demonstrate clonogenic potential and differentiate into cholangiocytes and functional hepatocytes in vitro, whereas colony formation by epithelial cell adhesion molecule-positive progenitor cells is blocked by αvβ6-neutralizing antibody and in integrin beta 6-deficient cells. Inhibition of progenitors by anti-αvβ6 antibody is recapitulated by TGFβ1 neutralization and rescued by addition of bioactive TGFβ1. Genetic disruption or selective targeting of αvβ6 with 3G9 antibody potently inhibits progenitor cell responses in mouse models of chronic biliary injury and protects from liver fibrosis and tumorigenesis, two conditions clinically associated with exacerbated ductular reaction. These results suggest that αvβ6 is a promising target for chronic fibrotic liver diseases and associated cancers.
Integrin αvβ6 is rapidly up-regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent transforming growth factor beta 1 (TGFβ1). In human liver cirrhosis, αvβ6 is overexpressed by cells comprising the ductular reaction, and its inhibition suppresses experimental biliary fibrosis in rodents. Here, we show that αvβ6 is expressed on the actively proliferating subset of hepatic progenitor cells and is required for their progenitor function in vivo and in vitro through integrin αvβ6-dependent TGFβ1 activation. Freshly isolated αvβ6+ liver cells demonstrate clonogenic potential and differentiate into cholangiocytes and functional hepatocytes in vitro, whereas colony formation by epithelial cell adhesion molecule-positive progenitor cells is blocked by αvβ6-neutralizing antibody and in integrin beta 6-deficient cells. Inhibition of progenitors by anti-αvβ6 antibody is recapitulated by TGFβ1 neutralization and rescued by addition of bioactive TGFβ1. Genetic disruption or selective targeting of αvβ6 with 3G9 antibody potently inhibits progenitor cell responses in mouse models of chronic biliary injury and protects from liver fibrosis and tumorigenesis, two conditions clinically associated with exacerbated ductular reaction.
UNLABELLEDIntegrin αvβ6 is rapidly up-regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent transforming growth factor beta 1 (TGFβ1). In human liver cirrhosis, αvβ6 is overexpressed by cells comprising the ductular reaction, and its inhibition suppresses experimental biliary fibrosis in rodents. Here, we show that αvβ6 is expressed on the actively proliferating subset of hepatic progenitor cells and is required for their progenitor function in vivo and in vitro through integrin αvβ6-dependent TGFβ1 activation. Freshly isolated αvβ6(+) liver cells demonstrate clonogenic potential and differentiate into cholangiocytes and functional hepatocytes in vitro, whereas colony formation by epithelial cell adhesion molecule-positive progenitor cells is blocked by αvβ6-neutralizing antibody and in integrin beta 6-deficient cells. Inhibition of progenitors by anti-αvβ6 antibody is recapitulated by TGFβ1 neutralization and rescued by addition of bioactive TGFβ1. Genetic disruption or selective targeting of αvβ6 with 3G9 antibody potently inhibits progenitor cell responses in mouse models of chronic biliary injury and protects from liver fibrosis and tumorigenesis, two conditions clinically associated with exacerbated ductular reaction.CONCLUSIONThese results suggest that αvβ6 is a promising target for chronic fibrotic liver diseases and associated cancers.
Integrin αvβ6 is rapidly up‐regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent transforming growth factor beta 1 (TGFβ1). In human liver cirrhosis, αvβ6 is overexpressed by cells comprising the ductular reaction, and its inhibition suppresses experimental biliary fibrosis in rodents. Here, we show that αvβ6 is expressed on the actively proliferating subset of hepatic progenitor cells and is required for their progenitor function in vivo and in vitro through integrin αvβ6‐dependent TGFβ1 activation. Freshly isolated αvβ6+ liver cells demonstrate clonogenic potential and differentiate into cholangiocytes and functional hepatocytes in vitro, whereas colony formation by epithelial cell adhesion molecule‐positive progenitor cells is blocked by αvβ6‐neutralizing antibody and in integrin beta 6‐deficient cells. Inhibition of progenitors by anti‐αvβ6 antibody is recapitulated by TGFβ1 neutralization and rescued by addition of bioactive TGFβ1. Genetic disruption or selective targeting of αvβ6 with 3G9 antibody potently inhibits progenitor cell responses in mouse models of chronic biliary injury and protects from liver fibrosis and tumorigenesis, two conditions clinically associated with exacerbated ductular reaction. Conclusion: These results suggest that αvβ6 is a promising target for chronic fibrotic liver diseases and associated cancers. (Hepatology 2016;63:217–232)
Integrin αvβ6 is rapidly up‐regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent transforming growth factor beta 1 (TGFβ1). In human liver cirrhosis, αvβ6 is overexpressed by cells comprising the ductular reaction, and its inhibition suppresses experimental biliary fibrosis in rodents. Here, we show that αvβ6 is expressed on the actively proliferating subset of hepatic progenitor cells and is required for their progenitor function in vivo and in vitro through integrin αvβ6‐dependent TGFβ1 activation. Freshly isolated αvβ6 + liver cells demonstrate clonogenic potential and differentiate into cholangiocytes and functional hepatocytes in vitro , whereas colony formation by epithelial cell adhesion molecule‐positive progenitor cells is blocked by αvβ6‐neutralizing antibody and in integrin beta 6‐deficient cells. Inhibition of progenitors by anti‐αvβ6 antibody is recapitulated by TGFβ1 neutralization and rescued by addition of bioactive TGFβ1. Genetic disruption or selective targeting of αvβ6 with 3G9 antibody potently inhibits progenitor cell responses in mouse models of chronic biliary injury and protects from liver fibrosis and tumorigenesis, two conditions clinically associated with exacerbated ductular reaction. Conclusion: These results suggest that αvβ6 is a promising target for chronic fibrotic liver diseases and associated cancers. (H epatology 2016;63:217–232)
Author Peng, Zhen‐Wei
Weinreb, Paul H.
Sheppard, Dean
Dixit, Richa
Schuppan, Detlef
Ikenaga, Naoki
Sverdlov, Deanna Y.
Popov, Yury
Liu, Susan B.
Violette, Shelia
Vaid, Kahini A.
AuthorAffiliation 1 Department of Oncology, The First Affiliated Hospital of Sun Yat-sen University, Guangzhou, China
2 Division of Gastroenterology and Hepatology, Beth Israel Deaconess Medical Center, Harvard Medical School, Boston, MA
5 Institute of Translational Immunology and Research Center for Immune Therapy, University Medical Center, Mainz, Germany
3 Biogen, Inc., Cambridge, MA
4 Lung Biology Center, University of California, San Francisco School of Medicine, San Francisco, CA
AuthorAffiliation_xml – name: 5 Institute of Translational Immunology and Research Center for Immune Therapy, University Medical Center, Mainz, Germany
– name: 3 Biogen, Inc., Cambridge, MA
– name: 2 Division of Gastroenterology and Hepatology, Beth Israel Deaconess Medical Center, Harvard Medical School, Boston, MA
– name: 1 Department of Oncology, The First Affiliated Hospital of Sun Yat-sen University, Guangzhou, China
– name: 4 Lung Biology Center, University of California, San Francisco School of Medicine, San Francisco, CA
Author_xml – sequence: 1
  givenname: Zhen‐Wei
  surname: Peng
  fullname: Peng, Zhen‐Wei
  organization: Harvard Medical School
– sequence: 2
  givenname: Naoki
  surname: Ikenaga
  fullname: Ikenaga, Naoki
  organization: Harvard Medical School
– sequence: 3
  givenname: Susan B.
  surname: Liu
  fullname: Liu, Susan B.
  organization: Harvard Medical School
– sequence: 4
  givenname: Deanna Y.
  surname: Sverdlov
  fullname: Sverdlov, Deanna Y.
  organization: Harvard Medical School
– sequence: 5
  givenname: Kahini A.
  surname: Vaid
  fullname: Vaid, Kahini A.
  organization: Harvard Medical School
– sequence: 6
  givenname: Richa
  surname: Dixit
  fullname: Dixit, Richa
  organization: Harvard Medical School
– sequence: 7
  givenname: Paul H.
  surname: Weinreb
  fullname: Weinreb, Paul H.
  organization: Biogen, Inc
– sequence: 8
  givenname: Shelia
  surname: Violette
  fullname: Violette, Shelia
  organization: Biogen, Inc
– sequence: 9
  givenname: Dean
  surname: Sheppard
  fullname: Sheppard, Dean
  organization: University of California, San Francisco School of Medicine
– sequence: 10
  givenname: Detlef
  surname: Schuppan
  fullname: Schuppan, Detlef
  organization: University Medical Center
– sequence: 11
  givenname: Yury
  surname: Popov
  fullname: Popov, Yury
  organization: Harvard Medical School
BackLink https://www.ncbi.nlm.nih.gov/pubmed/26448099$$D View this record in MEDLINE/PubMed
BookMark eNp9kd1qFTEUhYNU7Gn1wheQuVTotDt_83MjSKm2UNALvQ6ZTHIaySTHJFM5lz6SPkifycw5taigEAjs_e21NmsfoQMfvEboOYZTDEDObvTmlHSkZY_QCnPS1pRyOEArIC3UPab9ITpK6TMA9Ix0T9AhaRjroO9X6NuVz3odra_uvt_e_WgqFW22Sjq3raJez05mnapiIEu12sSw1t7mECulnavM7FW2wVfSj0tzCgs9ziqXwVgE5K59Uhk7xJBsOtmReZ5CtEVJl9JT9NhIl_Sz-_8YfXp78fH8sr5-_-7q_M11rRgAq6kiI8fQ9XSUIx1a3CnDJR047ThlsoeOADdMmpaVh83QQDO2zYAxmHEwmh6j13vdzTxMelTa5yid2EQ7ybgVQVrxZ8fbG7EOt4JTTICRIvDyXiCGL7NOWUw2LTFIr8OcBG45Zl3Zpinoi9-9Hkx-5V6AV3tAlVhS1OYBwSCWm4oSudjdtLBnf7HKZrnkWta07n8TX63T239Li8uLD_uJn2SCt_Q
CitedBy_id crossref_primary_10_1172_JCI186421
crossref_primary_10_1016_j_cgh_2024_10_005
crossref_primary_10_1016_j_jss_2017_03_051
crossref_primary_10_1016_j_jhep_2017_11_012
crossref_primary_10_1016_j_mucimm_2025_05_007
crossref_primary_10_1016_j_ejphar_2022_174866
crossref_primary_10_1055_s_0040_1705109
crossref_primary_10_1111_liv_15157
crossref_primary_10_3390_cancers11111783
crossref_primary_10_1016_j_biocel_2018_04_013
crossref_primary_10_1002_path_5680
crossref_primary_10_1172_JCI93561
crossref_primary_10_1053_j_gastro_2024_01_049
crossref_primary_10_1038_s41419_020_2372_9
crossref_primary_10_3390_ijms24076170
crossref_primary_10_1016_j_ajpath_2018_10_007
crossref_primary_10_1097_MOG_0000000000000809
crossref_primary_10_1002_hep4_1084
crossref_primary_10_1038_s41586_020_2938_9
crossref_primary_10_3389_fonc_2021_770997
crossref_primary_10_3390_livers5010005
crossref_primary_10_3390_ijms221910439
crossref_primary_10_3390_ijms19051294
crossref_primary_10_1016_j_ebiom_2018_08_024
crossref_primary_10_1126_science_adp0246
crossref_primary_10_1097_HEP_0000000000000026
crossref_primary_10_3389_fimmu_2023_1192840
crossref_primary_10_12688_f1000research_14578_1
crossref_primary_10_1016_j_lfs_2020_118354
crossref_primary_10_3390_ijms19124115
crossref_primary_10_3390_pharmaceutics14010217
crossref_primary_10_1016_j_bbadis_2017_07_024
crossref_primary_10_3389_fnut_2022_1032722
crossref_primary_10_1016_j_gene_2019_04_078
crossref_primary_10_1038_s41419_025_07590_4
crossref_primary_10_1016_j_coph_2021_11_003
crossref_primary_10_1016_j_bcp_2023_115740
crossref_primary_10_1016_j_gtc_2017_01_011
crossref_primary_10_1038_s41575_020_00372_7
crossref_primary_10_1172_JCI132305
crossref_primary_10_1016_j_jhep_2020_12_010
crossref_primary_10_1016_j_bbadis_2017_06_027
crossref_primary_10_1080_0886022X_2024_2409348
crossref_primary_10_1042_BCJ20200749
crossref_primary_10_1053_j_gastro_2024_07_044
crossref_primary_10_33549_physiolres_933499
crossref_primary_10_1016_j_matbio_2018_04_006
crossref_primary_10_1097_HC9_0000000000000140
crossref_primary_10_3389_fphar_2025_1517491
crossref_primary_10_1016_j_lfs_2022_120882
crossref_primary_10_3389_fchem_2021_674786
crossref_primary_10_1002_path_5228
crossref_primary_10_1016_j_jhep_2016_10_033
crossref_primary_10_3390_ijms19123875
crossref_primary_10_1016_j_mam_2018_09_002
crossref_primary_10_3390_biom11081095
crossref_primary_10_1007_s41048_018_0071_1
crossref_primary_10_1016_j_addr_2017_07_014
crossref_primary_10_1007_s11926_018_0710_z
crossref_primary_10_1111_1751_2980_12879
crossref_primary_10_1111_liv_14490
crossref_primary_10_3390_cells8101279
crossref_primary_10_1093_toxsci_kfac128
crossref_primary_10_1007_s43152_020_00004_x
crossref_primary_10_1016_j_jbc_2021_100657
crossref_primary_10_1002_hep_30550
crossref_primary_10_1016_j_ajpath_2024_01_023
crossref_primary_10_1016_j_bbadis_2021_166290
crossref_primary_10_1177_15330338231189399
crossref_primary_10_1080_14656566_2024_2362263
crossref_primary_10_1097_MOG_0000000000000995
crossref_primary_10_1007_s11302_019_09664_3
crossref_primary_10_1038_s41584_019_0324_5
crossref_primary_10_3390_cells8121503
crossref_primary_10_1055_a_2601_9426
crossref_primary_10_1371_journal_pone_0292998
crossref_primary_10_1007_s12026_025_09622_9
crossref_primary_10_1016_j_matbio_2017_11_002
crossref_primary_10_1016_j_bbadis_2018_07_025
crossref_primary_10_3389_fimmu_2022_866040
crossref_primary_10_3390_ijms25147873
crossref_primary_10_1002_path_6002
crossref_primary_10_12688_f1000research_8827_1
crossref_primary_10_1007_s00441_016_2407_9
crossref_primary_10_1093_ecco_jcc_jjae131
crossref_primary_10_1111_cas_14455
crossref_primary_10_1016_j_jhep_2020_05_049
crossref_primary_10_1016_j_cellimm_2017_08_004
crossref_primary_10_1016_j_yexcr_2021_113003
crossref_primary_10_1016_j_mtbio_2025_102031
crossref_primary_10_3390_cells9092027
crossref_primary_10_14814_phy2_14508
crossref_primary_10_3390_cancers11050615
crossref_primary_10_1126_scitranslmed_aaw2050
Cites_doi 10.1038/nature11826
10.1038/nm1377
10.1016/j.ccr.2009.08.021
10.1002/hep.21466
10.1172/JCI23486
10.1053/gast.1996.v111.pm8698195
10.1053/j.gastro.2011.01.040
10.1055/s-2004-815564
10.1053/j.gastro.2008.04.009
10.1002/hep.21849
10.1111/j.1365-2559.2006.02468.x
10.1016/S0140-6736(05)67530-7
10.2353/ajpath.2007.061133
10.1016/j.ajpath.2014.02.020
10.1101/gad.2029411
10.1074/jbc.M312103200
10.1016/j.jhep.2005.06.025
10.1038/nature10152
10.1111/j.1365-2559.2004.01919.x
10.1016/j.ajpath.2013.03.018
10.1002/hep.27659
10.1038/nrgastro.2012.23
10.1002/hep.26753
10.1126/science.1118389
10.1016/j.ajpath.2014.10.013
10.1002/hep.20650
10.1016/S0092-8674(00)80545-0
10.1016/j.jhep.2007.11.021
10.1016/j.stem.2014.06.003
10.1016/j.ccr.2008.03.007
10.1038/nbt.2464
10.1074/jbc.271.42.25976
10.1016/j.stem.2014.09.008
10.1002/hep.23123
10.1016/j.jhep.2005.06.015
10.1016/j.jhep.2009.12.006
10.1002/hep.27425
10.1053/j.gastro.2007.05.012
10.1002/hep.22238
10.1177/41.10.8245410
10.1053/j.gastro.2004.08.006
10.1038/nature01413
10.1101/gad.2027811
10.1083/jcb.133.4.921
10.1053/j.gastro.2014.08.038
10.1242/dev.031369
ContentType Journal Article
Copyright 2015 by the American Association for the Study of Liver Diseases
2015 by the American Association for the Study of Liver Diseases.
Copyright_xml – notice: 2015 by the American Association for the Study of Liver Diseases
– notice: 2015 by the American Association for the Study of Liver Diseases.
DBID AAYXX
CITATION
CGR
CUY
CVF
ECM
EIF
NPM
7U8
7X8
C1K
JXQ
5PM
DOI 10.1002/hep.28274
DatabaseName CrossRef
Medline
MEDLINE
MEDLINE (Ovid)
MEDLINE
MEDLINE
PubMed
TOXLINE
MEDLINE - Academic
Environmental Sciences and Pollution Management
Toxline
PubMed Central (Full Participant titles)
DatabaseTitle CrossRef
MEDLINE
Medline Complete
MEDLINE with Full Text
PubMed
MEDLINE (Ovid)
TOXLINE
MEDLINE - Academic
Environmental Sciences and Pollution Management
DatabaseTitleList MEDLINE

TOXLINE

CrossRef
Database_xml – sequence: 1
  dbid: NPM
  name: PubMed
  url: http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?db=PubMed
  sourceTypes: Index Database
– sequence: 2
  dbid: 7X8
  name: MEDLINE - Academic
  url: https://search.proquest.com/medline
  sourceTypes: Aggregation Database
DeliveryMethod fulltext_linktorsrc
Discipline Medicine
EISSN 1527-3350
EndPage 232
ExternalDocumentID PMC5312042
26448099
10_1002_hep_28274
HEP28274
Genre article
Research Support, Non-U.S. Gov't
Journal Article
GrantInformation_xml – fundername: NIDDK NIH HHS
  grantid: P30 DK026743
GroupedDBID ---
--K
.3N
.55
.GA
.GJ
.Y3
05W
0R~
10A
186
1B1
1CY
1L6
1OC
1ZS
1~5
31~
33P
3O-
3SF
3WU
4.4
4G.
4ZD
50Y
50Z
51W
51X
52M
52N
52O
52P
52R
52S
52T
52U
52V
52W
52X
53G
5GY
5RE
5VS
7-5
702
7PT
8-0
8-1
8-3
8-4
8-5
8UM
930
A01
A03
AAEDT
AAESR
AAEVG
AAHHS
AALRI
AANHP
AAONW
AAQFI
AAQQT
AAQXK
AASGY
AAXRX
AAXUO
AAZKR
ABCQN
ABCUV
ABEML
ABIJN
ABJNI
ABLJU
ABMAC
ABOCM
ABPVW
ABWVN
ABXGK
ACAHQ
ACBWZ
ACCFJ
ACCZN
ACGFS
ACLDA
ACMXC
ACPOU
ACPRK
ACRPL
ACSCC
ACXBN
ACXQS
ACYXJ
ACZKN
ADBBV
ADEOM
ADIZJ
ADKYN
ADMGS
ADMUD
ADNMO
ADOZA
ADXAS
ADZMN
ADZOD
AECAP
AEEZP
AEIMD
AENEX
AEQDE
AFBPY
AFFNX
AFGKR
AFNMH
AFUWQ
AFZJQ
AGQPQ
AHMBA
AHQVU
AIACR
AIURR
AIWBW
AJAOE
AJBDE
ALAGY
ALMA_UNASSIGNED_HOLDINGS
ALUQN
AMBMR
AMYDB
ASPBG
ATUGU
AVWKF
AZBYB
AZFZN
AZVAB
BAFTC
BAWUL
BDRZF
BHBCM
BMXJE
BROTX
BRXPI
BY8
C45
CAG
COF
CS3
D-6
D-7
D-E
D-F
DCZOG
DIK
DPXWK
DR2
DRFUL
DRMAN
DRSTM
DU5
E3Z
EBS
EJD
F00
F01
F04
F5P
FD8
FDB
FEDTE
FGOYB
FUBAC
G-S
G.N
GNP
GODZA
H.X
HBH
HF~
HHY
HHZ
HVGLF
HZ~
IHE
IX1
J0M
J5H
JPC
KBYEO
KQQ
LATKE
LC2
LC3
LEEKS
LH4
LITHE
LOXES
LP6
LP7
LUTES
LW6
LYRES
M41
M65
MEWTI
MJL
MK4
MRFUL
MRMAN
MRSTM
MSFUL
MSMAN
MSSTM
MXFUL
MXMAN
MXSTM
N04
N05
N4W
N9A
NF~
NNB
NQ-
O66
O9-
OIG
OK1
OVD
P2P
P2W
P2X
P2Z
P4B
P4D
PALCI
PQQKQ
Q.N
Q11
QB0
QRW
R.K
R2-
RIG
RIWAO
RJQFR
ROL
RPZ
RX1
RYL
SEW
SSZ
SUPJJ
TEORI
UB1
V2E
V9Y
W2D
W8V
W99
WBKPD
WH7
WHWMO
WIB
WIH
WIJ
WIK
WJL
WOHZO
WQJ
WVDHM
WXI
WXSBR
X7M
XG1
XV2
ZGI
ZXP
ZZTAW
~IA
~WT
9DU
AAMMB
AAYXX
ADSXY
AEFGJ
AGXDD
AIDQK
AIDYY
AIQQE
CITATION
O8X
CGR
CUY
CVF
ECM
EIF
NPM
7U8
7X8
C1K
JXQ
5PM
ID FETCH-LOGICAL-c4004-3c2d510893dad3b718cf5a3b538534a908205f4af74f741fb606d76b110fdbfe3
IEDL.DBID DRFUL
ISSN 0270-9139
IngestDate Tue Sep 30 16:45:55 EDT 2025
Fri Sep 05 10:58:04 EDT 2025
Mon Jul 21 05:51:37 EDT 2025
Sat Nov 29 05:40:36 EST 2025
Tue Nov 18 19:51:19 EST 2025
Thu Apr 24 03:18:00 EDT 2025
IsDoiOpenAccess false
IsOpenAccess true
IsPeerReviewed true
IsScholarly true
Issue 1
Language English
License 2015 by the American Association for the Study of Liver Diseases.
LinkModel DirectLink
MergedId FETCHMERGED-LOGICAL-c4004-3c2d510893dad3b718cf5a3b538534a908205f4af74f741fb606d76b110fdbfe3
Notes Potential conflict of interest: Dr. Sheppard consults and owns stock in Pliant Therapeutics. He holds intellectual property rights and received grants from Biogen. He advises Genentech. Dr. Schuppan consults and received grants from Boehringer‐Ingelheim. He consults for MSD, Mitsubishi‐Tanabe, and Takeda. Dr. Popov received grants from Biogen and Stromedix. Dr. Violette is employed by and owns stock in Biogen. Dr. Weinreb is employed by and owns stock in Biogen.
Supported by research grants from Biogen, Inc., and Stomedix, Inc.; an institutional grant from Department of Medicine, Beth Israel Deaconess Medical Center (to Y.P.); a career development award from The First Affiliated Hospital of Sun Yat‐sen University (to Z.‐W.P.); and grants from the National Natural Science Foundation of China (81301842) and the Pearl River S&T Nova Program (2014J2200087).
ObjectType-Article-1
SourceType-Scholarly Journals-1
ObjectType-Feature-2
content type line 23
OpenAccessLink https://aasldpubs.onlinelibrary.wiley.com/doi/pdfdirect/10.1002/hep.28274
PMID 26448099
PQID 1751488536
PQPubID 23479
PageCount 16
ParticipantIDs pubmedcentral_primary_oai_pubmedcentral_nih_gov_5312042
proquest_miscellaneous_1751488536
pubmed_primary_26448099
crossref_primary_10_1002_hep_28274
crossref_citationtrail_10_1002_hep_28274
wiley_primary_10_1002_hep_28274_HEP28274
PublicationCentury 2000
PublicationDate January 2016
PublicationDateYYYYMMDD 2016-01-01
PublicationDate_xml – month: 01
  year: 2016
  text: January 2016
PublicationDecade 2010
PublicationPlace United States
PublicationPlace_xml – name: United States
PublicationTitle Hepatology (Baltimore, Md.)
PublicationTitleAlternate Hepatology
PublicationYear 2016
References 2015; 185
2004; 127
2006; 12
2005; 310
2005; 115
2004; 45
1993; 41
2005; 41
2005; 43
2008; 13
2013; 183
2009; 136
2011; 474
1998; 15
2009; 51
2004; 279
2009; 50
2005; 366
2015; 61
2007; 171
2006; 44
2006; 49
2007; 133
1995; 108
2013; 31
2008; 47
2008; 48
2014; 15
2014; 59
1996; 271
1996; 133
1996; 111
1999; 96
2014; 184
2008; 135
2011; 25
2013; 494
2003; 422
2011; 140
2007; 45
2009; 16
2010; 52
2007; 46
2014; 147
2003; 100
2012; 9
2003; 23
(hep28274-bib-0039-20241017) 2006; 49
(hep28274-bib-0029-20241017) 2011; 25
(hep28274-bib-0036-20241017) 2012; 9
(hep28274-bib-0020-20241017) 2005; 43
(hep28274-bib-0018-20241017) 2004; 45
(hep28274-bib-0023-20241017) 1996; 133
(hep28274-bib-0006-20241017) 2008; 47
(hep28274-bib-0033-20241017) 2009; 136
(hep28274-bib-0002-20241017) 2005; 366
(hep28274-bib-0045-20241017) 2005; 310
(hep28274-bib-0031-20241017) 2013; 183
(hep28274-bib-0041-20241017) 2005; 41
(hep28274-bib-0013-20241017) 1999; 96
(hep28274-bib-0010-20241017) 1995; 108
(hep28274-bib-0050-20241017) 2006; 44
(hep28274-bib-0047-20241017) 2015; 61
(hep28274-bib-0024-20241017) 2003; 422
(hep28274-bib-0043-20241017) 1996; 271
(hep28274-bib-0011-20241017) 1993; 41
(hep28274-bib-0038-20241017) 2003; 100
(hep28274-bib-0028-20241017) 2008; 13
(hep28274-bib-0040-20241017) 2003; 23
(hep28274-bib-0044-20241017) 2007; 45
(hep28274-bib-0016-20241017) 2008; 135
(hep28274-bib-0025-20241017) 1996; 111
(hep28274-bib-0009-20241017) 2009; 16
(hep28274-bib-0005-20241017) 2009; 50
(hep28274-bib-0026-20241017) 2004; 279
(hep28274-bib-0008-20241017) 2005; 115
(hep28274-bib-0001-20241017) 2009; 51
(hep28274-bib-0042-20241017) 2007; 133
(hep28274-bib-0032-20241017) 2015; 185
(hep28274-bib-0030-20241017) 2011; 140
(hep28274-bib-0003-20241017) 1998; 15
(hep28274-bib-0046-20241017) 2014; 184
(hep28274-bib-0014-20241017) 2008; 48
(hep28274-bib-0017-20241017) 2007; 46
(hep28274-bib-0019-20241017) 2015; 61
(hep28274-bib-0027-20241017) 2014; 147
(hep28274-bib-0007-20241017) 2006; 12
(hep28274-bib-0037-20241017) 2014; 59
(hep28274-bib-0012-20241017) 2011; 474
(hep28274-bib-0034-20241017) 2014; 15
(hep28274-bib-0035-20241017) 2014; 15
(hep28274-bib-0049-20241017) 2013; 494
(hep28274-bib-0015-20241017) 2010; 52
(hep28274-bib-0021-20241017) 2007; 171
(hep28274-bib-0004-20241017) 2004; 127
(hep28274-bib-0048-20241017) 2011; 25
(hep28274-bib-0022-20241017) 2013; 31
12902545 - Proc Natl Acad Sci U S A. 2003 Sep 30;100 Suppl 1:11881-8
23680654 - Am J Pathol. 2013 Jul;183(1):182-94
19711424 - Hepatology. 2009 Oct;50(4):1294-306
17187411 - Hepatology. 2007 Jan;45(1):31-41
21632826 - Genes Dev. 2011 Jun 1;25(11):1193-203
25503476 - Hepatology. 2015 Jun;61(6):2100-3
22371217 - Nat Rev Gastroenterol Hepatol. 2012 Feb 28;9(4):231-40
18393293 - Hepatology. 2008 May;47(5):1544-56
17600122 - Am J Pathol. 2007 Aug;171(2):525-36
18455123 - Cancer Cell. 2008 May;13(5):394-406
19800575 - Cancer Cell. 2009 Oct 6;16(4):295-308
25312494 - Cell Stem Cell. 2014 Nov 6;15(5):605-18
15521023 - Gastroenterology. 2004 Nov;127(5):1565-77
16532004 - Nat Med. 2006 Apr;12(4):410-6
20137822 - J Hepatol. 2010 Mar;52(3):362-9
17631134 - Gastroenterology. 2007 Jul;133(1):80-90
16210527 - Science. 2005 Oct 7;310(5745):68-71
21632825 - Genes Dev. 2011 Jun 1;25(11):1185-92
8824234 - J Biol Chem. 1996 Oct 18;271(42):25976-80
25478810 - Am J Pathol. 2015 Feb;185(2):325-34
20101749 - Hepatology. 2010 Feb;51(2):660-78
16110324 - J Clin Invest. 2005 Sep;115(9):2330-40
16223543 - J Hepatol. 2005 Dec;43(6):1045-54
23242163 - Nat Biotechnol. 2013 Jan;31(1):63-70
14960589 - J Biol Chem. 2004 Apr 23;279(17 ):17875-87
24115180 - Hepatology. 2014 Apr;59(4):1617-26
16168515 - J Hepatol. 2006 Jan;44(1):167-75
8666675 - J Cell Biol. 1996 May;133(4):921-8
9845427 - Semin Diagn Pathol. 1998 Nov;15(4):259-69
10025398 - Cell. 1999 Feb 5;96(3):319-28
15330800 - Histopathology. 2004 Sep;45(3):226-36
19429791 - Development. 2009 Jun;136(11):1951-60
15793848 - Hepatology. 2005 Apr;41(4):809-18
18538673 - Gastroenterology. 2008 Aug;135(2):660-70
25130492 - Cell Stem Cell. 2014 Sep 4;15(3):340-9
17924447 - Hepatology. 2007 Nov;46(5):1404-12
25203810 - Hepatology. 2015 Feb;61(2):678-91
24681248 - Am J Pathol. 2014 May;184(5):1263-7
25173753 - Gastroenterology. 2014 Dec;147(6):1378-92
8245410 - J Histochem Cytochem. 1993 Oct;41(10):1521-7
14722815 - Semin Liver Dis. 2003 Nov;23(4):385-96
18221819 - J Hepatol. 2008 Mar;48(3):453-64
21677751 - Nature. 2011 Jun 15;474(7351):343-9
12634787 - Nature. 2003 Mar 13;422(6928):169-73
21277850 - Gastroenterology. 2011 May;140(5):1642-52
23354049 - Nature. 2013 Feb 14;494(7436):247-50
8698195 - Gastroenterology. 1996 Jul;111(1):165-71
7673344 - J Cell Sci. 1995 Jun;108 ( Pt 6):2241-51
16879391 - Histopathology. 2006 Aug;49(2):138-51
16214602 - Lancet. 2005 Oct 8;366(9493):1303-14
References_xml – volume: 184
  start-page: 1263
  year: 2014
  end-page: 1267
  article-title: The liver is a peculiar organ when it comes to stem cells
  publication-title: Am J Pathol
– volume: 41
  start-page: 1521
  year: 1993
  end-page: 1527
  article-title: Restricted distribution of integrin beta 6 mRNA in primate epithelial tissues
  publication-title: J Histochem Cytochem
– volume: 49
  start-page: 138
  year: 2006
  end-page: 151
  article-title: The clinicopathological and prognostic relevance of cytokeratin 7 and 19 expression in hepatocellular carcinoma. A possible progenitor cell origin
  publication-title: Histopathology
– volume: 115
  start-page: 2330
  year: 2005
  end-page: 2340
  article-title: TWEAK induces liver progenitor cell proliferation
  publication-title: J Clin Invest
– volume: 15
  start-page: 259
  year: 1998
  end-page: 269
  article-title: Ductular reaction and its diagnostic significance
  publication-title: Semin Diagn Pathol
– volume: 96
  start-page: 319
  year: 1999
  end-page: 328
  article-title: The integrin alphavbeta6 binds and activates latent TGF beta 1: a mechanism for regulating pulmonary inflammation and fibrosis
  publication-title: Cell
– volume: 135
  start-page: 660
  year: 2008
  end-page: 670
  article-title: Inhibition of integrin alphavbeta6 on cholangiocytes blocks transforming growth factor‐beta activation and retards biliary fibrosis progression
  publication-title: Gastroenterology
– volume: 41
  start-page: 809
  year: 2005
  end-page: 818
  article-title: Fibrosis correlates with a ductular reaction in hepatitis C: roles of impaired replication, progenitor cells and steatosis
  publication-title: Hepatology
– volume: 12
  start-page: 410
  year: 2006
  end-page: 416
  article-title: A novel prognostic subtype of human hepatocellular carcinoma derived from hepatic progenitor cells
  publication-title: Nat Med
– volume: 140
  start-page: 1642
  year: 2011
  end-page: 1652
  article-title: Tissue transglutaminase does not affect fibrotic matrix stability or regression of liver fibrosis in mice
  publication-title: Gastroenterology
– volume: 271
  start-page: 25976
  year: 1996
  end-page: 25980
  article-title: Identification of a sequence within the integrin beta6 subunit cytoplasmic domain that is required to support the specific effect of alphavbeta6 on proliferation in three‐dimensional culture
  publication-title: J Biol Chem
– volume: 52
  start-page: 362
  year: 2010
  end-page: 369
  article-title: The alphavbeta6 integrin is a highly specific immunohistochemical marker for cholangiocarcinoma
  publication-title: J Hepatol
– volume: 366
  start-page: 1303
  year: 2005
  end-page: 1314
  article-title: Cholangiocarcinoma
  publication-title: Lancet
– volume: 13
  start-page: 394
  year: 2008
  end-page: 406
  article-title: Regulation of to invasive breast carcinoma transition
  publication-title: Cancer Cell
– volume: 310
  start-page: 68
  year: 2005
  end-page: 71
  article-title: Transforming growth factor‐beta signaling in stem cells and cancer
  publication-title: Science
– volume: 59
  start-page: 1617
  year: 2014
  end-page: 1626
  article-title: Liver regeneration by stem/progenitor cells
  publication-title: Hepatology
– volume: 111
  start-page: 165
  year: 1996
  end-page: 171
  article-title: Effects of ursodeoxycholate and cholate feeding on liver disease in FVB mice with a disrupted mdr2 P‐glycoprotein gene
  publication-title: Gastroenterology
– volume: 422
  start-page: 169
  year: 2003
  end-page: 173
  article-title: Loss of integrin alpha(v)beta6‐mediated TGF‐beta activation causes Mmp12‐dependent emphysema
  publication-title: Nature
– volume: 61
  start-page: 2100
  year: 2015
  end-page: 2103
  article-title: Determining the fate of hepatic cells by lineage tracing: facts and pitfalls
  publication-title: Hepatology
– volume: 25
  start-page: 1185
  year: 2011
  end-page: 1192
  article-title: Foxl1‐Cre‐marked adult hepatic progenitors have clonogenic and bilineage differentiation potential
  publication-title: Genes Dev
– volume: 61
  start-page: 678
  year: 2015
  end-page: 691
  article-title: Connective tissue growth factor and integrin alphavbeta6: a new pair of regulators critical for ductular reaction and biliary fibrosis in mice
  publication-title: Hepatology
– volume: 31
  start-page: 63
  year: 2013
  end-page: 70
  article-title: Mass‐encoded synthetic biomarkers for multiplexed urinary monitoring of disease
  publication-title: Nat Biotechnol
– volume: 171
  start-page: 525
  year: 2007
  end-page: 536
  article-title: A new xenobiotic‐induced mouse model of sclerosing cholangitis and biliary fibrosis
  publication-title: Am J Pathol
– volume: 133
  start-page: 80
  year: 2007
  end-page: 90
  article-title: Progressive fibrosis in nonalcoholic steatohepatitis: association with altered regeneration and a ductular reaction
  publication-title: Gastroenterology
– volume: 47
  start-page: 1544
  year: 2008
  end-page: 1556
  article-title: Clinicopathological study on cholangiolocellular carcinoma suggesting hepatic progenitor cell origin
  publication-title: Hepatology
– volume: 147
  start-page: 1378
  year: 2014
  end-page: 1392
  article-title: Extrahepatic platelet‐derived growth factor‐beta, delivered by platelets, promotes activation of hepatic stellate cells and biliary fibrosis in mice
  publication-title: Gastroenterology
– volume: 108
  start-page: 2241
  issue: Pt. 6
  year: 1995
  end-page: 2251
  article-title: Expression of the beta 6 integrin subunit in development, neoplasia and tissue repair suggests a role in epithelial remodeling
  publication-title: J Cell Sci
– volume: 25
  start-page: 1193
  year: 2011
  end-page: 1203
  article-title: Prospective isolation of a bipotential clonogenic liver progenitor cell in adult mice
  publication-title: Genes Dev
– volume: 48
  start-page: 453
  year: 2008
  end-page: 464
  article-title: Integrin alphavbeta6 is a marker of the progression of biliary and portal liver fibrosis and a novel target for antifibrotic therapies
  publication-title: J Hepatol
– volume: 133
  start-page: 921
  year: 1996
  end-page: 928
  article-title: Inactivation of the integrin beta6 subunit gene reveals a role of epithelial integrins in regulating inflammation in the lung and skin
  publication-title: J Cell Biol
– volume: 46
  start-page: 1404
  year: 2007
  end-page: 1412
  article-title: Role of alphavbeta6 integrin in acute biliary fibrosis
  publication-title: Hepatology
– volume: 23
  start-page: 385
  year: 2003
  end-page: 396
  article-title: Progenitor cells in diseased human liver
  publication-title: Semin Liver Dis
– volume: 51
  start-page: 660
  year: 2009
  end-page: 678
  article-title: Diagnosis and management of primary sclerosing cholangitis
  publication-title: Hepatology
– volume: 183
  start-page: 182
  year: 2013
  end-page: 194
  article-title: Failure of fibrotic liver regeneration in mice is linked to a severe fibrogenic response driven by hepatic progenitor cell activation
  publication-title: Am J Pathol
– volume: 136
  start-page: 1951
  year: 2009
  end-page: 1960
  article-title: Potential hepatic stem cells reside in EpCAM cells of normal and injured mouse liver
  publication-title: Development
– volume: 45
  start-page: 226
  year: 2004
  end-page: 236
  article-title: Immunohistochemical screening for beta6‐integrin subunit expression in adenocarcinomas using a novel monoclonal antibody reveals strong up‐regulation in pancreatic ductal adenocarcinomas and
  publication-title: Histopathology
– volume: 44
  start-page: 167
  year: 2006
  end-page: 175
  article-title: Oral imatinib treatment reduces early fibrogenesis but does not prevent progression in the long term
  publication-title: J Hepatol
– volume: 9
  start-page: 231
  year: 2012
  end-page: 240
  article-title: The biliary tree—a reservoir of multipotent stem cells
  publication-title: Nat Rev Gastroenterol Hepatol
– volume: 15
  start-page: 340
  year: 2014
  end-page: 349
  article-title: Adult hepatocytes are generated by self‐duplication rather than stem cell differentiation
  publication-title: Cell Stem Cell
– volume: 100
  start-page: 11881
  issue: Suppl. 1
  year: 2003
  end-page: 11888
  article-title: The origin and liver repopulating capacity of murine oval cells
  publication-title: Proc Natl Acad Sci USA
– volume: 50
  start-page: 1294
  year: 2009
  end-page: 1306
  article-title: Targeting liver fibrosis: strategies for development and validation of antifibrotic therapies
  publication-title: Hepatology
– volume: 494
  start-page: 247
  year: 2013
  end-page: 250
  article-title: expansion of single Lgr5 liver stem cells induced by Wnt‐driven regeneration
  publication-title: Nature
– volume: 185
  start-page: 325
  year: 2015
  end-page: 334
  article-title: A new mdr2 mouse model of sclerosing cholangitis with rapid fibrosis progression, early‐onset portal hypertension, and liver cancer
  publication-title: Am J Pathol
– volume: 16
  start-page: 295
  year: 2009
  end-page: 308
  article-title: A lymphotoxin‐driven pathway to hepatocellular carcinoma
  publication-title: Cancer Cell
– volume: 45
  start-page: 31
  year: 2007
  end-page: 41
  article-title: Transforming growth factor‐beta differentially regulates oval cell and hepatocyte proliferation
  publication-title: Hepatology
– volume: 43
  start-page: 1045
  year: 2005
  end-page: 1054
  article-title: Mdr2 (Abcb4) mice spontaneously develop severe biliary fibrosis via massive dysregulation of pro‐ and antifibrogenic genes
  publication-title: J Hepatol
– volume: 474
  start-page: 343
  year: 2011
  end-page: 349
  article-title: Latent TGF‐beta structure and activation
  publication-title: Nature
– volume: 279
  start-page: 17875
  year: 2004
  end-page: 17887
  article-title: Function‐blocking integrin alphavbeta6 monoclonal antibodies: distinct ligand‐mimetic and nonligand‐mimetic classes
  publication-title: J Biol Chem
– volume: 15
  start-page: 605
  year: 2014
  end-page: 618
  article-title: Bipotential adult liver progenitors are derived from chronically injured mature hepatocytes
  publication-title: Cell Stem Cell
– volume: 127
  start-page: 1565
  year: 2004
  end-page: 1577
  article-title: The cholangiopathies: disorders of biliary epithelia
  publication-title: Gastroenterology
– volume: 108
  start-page: 2241
  issue: Pt. 6
  year: 1995
  ident: hep28274-bib-0010-20241017
  article-title: Expression of the beta 6 integrin subunit in development, neoplasia and tissue repair suggests a role in epithelial remodeling
  publication-title: J Cell Sci
– volume: 494
  start-page: 247
  year: 2013
  ident: hep28274-bib-0049-20241017
  article-title: In vitro expansion of single Lgr5+ liver stem cells induced by Wnt‐driven regeneration
  publication-title: Nature
  doi: 10.1038/nature11826
– volume: 12
  start-page: 410
  year: 2006
  ident: hep28274-bib-0007-20241017
  article-title: A novel prognostic subtype of human hepatocellular carcinoma derived from hepatic progenitor cells
  publication-title: Nat Med
  doi: 10.1038/nm1377
– volume: 16
  start-page: 295
  year: 2009
  ident: hep28274-bib-0009-20241017
  article-title: A lymphotoxin‐driven pathway to hepatocellular carcinoma
  publication-title: Cancer Cell
  doi: 10.1016/j.ccr.2009.08.021
– volume: 45
  start-page: 31
  year: 2007
  ident: hep28274-bib-0044-20241017
  article-title: Transforming growth factor‐beta differentially regulates oval cell and hepatocyte proliferation
  publication-title: Hepatology
  doi: 10.1002/hep.21466
– volume: 115
  start-page: 2330
  year: 2005
  ident: hep28274-bib-0008-20241017
  article-title: TWEAK induces liver progenitor cell proliferation
  publication-title: J Clin Invest
  doi: 10.1172/JCI23486
– volume: 111
  start-page: 165
  year: 1996
  ident: hep28274-bib-0025-20241017
  article-title: Effects of ursodeoxycholate and cholate feeding on liver disease in FVB mice with a disrupted mdr2 P‐glycoprotein gene
  publication-title: Gastroenterology
  doi: 10.1053/gast.1996.v111.pm8698195
– volume: 140
  start-page: 1642
  year: 2011
  ident: hep28274-bib-0030-20241017
  article-title: Tissue transglutaminase does not affect fibrotic matrix stability or regression of liver fibrosis in mice
  publication-title: Gastroenterology
  doi: 10.1053/j.gastro.2011.01.040
– volume: 23
  start-page: 385
  year: 2003
  ident: hep28274-bib-0040-20241017
  article-title: Progenitor cells in diseased human liver
  publication-title: Semin Liver Dis
  doi: 10.1055/s-2004-815564
– volume: 135
  start-page: 660
  year: 2008
  ident: hep28274-bib-0016-20241017
  article-title: Inhibition of integrin alphavbeta6 on cholangiocytes blocks transforming growth factor‐beta activation and retards biliary fibrosis progression
  publication-title: Gastroenterology
  doi: 10.1053/j.gastro.2008.04.009
– volume: 46
  start-page: 1404
  year: 2007
  ident: hep28274-bib-0017-20241017
  article-title: Role of alphavbeta6 integrin in acute biliary fibrosis
  publication-title: Hepatology
  doi: 10.1002/hep.21849
– volume: 49
  start-page: 138
  year: 2006
  ident: hep28274-bib-0039-20241017
  article-title: The clinicopathological and prognostic relevance of cytokeratin 7 and 19 expression in hepatocellular carcinoma. A possible progenitor cell origin
  publication-title: Histopathology
  doi: 10.1111/j.1365-2559.2006.02468.x
– volume: 366
  start-page: 1303
  year: 2005
  ident: hep28274-bib-0002-20241017
  article-title: Cholangiocarcinoma
  publication-title: Lancet
  doi: 10.1016/S0140-6736(05)67530-7
– volume: 171
  start-page: 525
  year: 2007
  ident: hep28274-bib-0021-20241017
  article-title: A new xenobiotic‐induced mouse model of sclerosing cholangitis and biliary fibrosis
  publication-title: Am J Pathol
  doi: 10.2353/ajpath.2007.061133
– volume: 184
  start-page: 1263
  year: 2014
  ident: hep28274-bib-0046-20241017
  article-title: The liver is a peculiar organ when it comes to stem cells
  publication-title: Am J Pathol
  doi: 10.1016/j.ajpath.2014.02.020
– volume: 25
  start-page: 1193
  year: 2011
  ident: hep28274-bib-0029-20241017
  article-title: Prospective isolation of a bipotential clonogenic liver progenitor cell in adult mice
  publication-title: Genes Dev
  doi: 10.1101/gad.2029411
– volume: 279
  start-page: 17875
  year: 2004
  ident: hep28274-bib-0026-20241017
  article-title: Function‐blocking integrin alphavbeta6 monoclonal antibodies: distinct ligand‐mimetic and nonligand‐mimetic classes
  publication-title: J Biol Chem
  doi: 10.1074/jbc.M312103200
– volume: 43
  start-page: 1045
  year: 2005
  ident: hep28274-bib-0020-20241017
  article-title: Mdr2 (Abcb4)−/− mice spontaneously develop severe biliary fibrosis via massive dysregulation of pro‐ and antifibrogenic genes
  publication-title: J Hepatol
  doi: 10.1016/j.jhep.2005.06.025
– volume: 474
  start-page: 343
  year: 2011
  ident: hep28274-bib-0012-20241017
  article-title: Latent TGF‐beta structure and activation
  publication-title: Nature
  doi: 10.1038/nature10152
– volume: 45
  start-page: 226
  year: 2004
  ident: hep28274-bib-0018-20241017
  article-title: Immunohistochemical screening for beta6‐integrin subunit expression in adenocarcinomas using a novel monoclonal antibody reveals strong up‐regulation in pancreatic ductal adenocarcinomas in vivo and in vitro
  publication-title: Histopathology
  doi: 10.1111/j.1365-2559.2004.01919.x
– volume: 183
  start-page: 182
  year: 2013
  ident: hep28274-bib-0031-20241017
  article-title: Failure of fibrotic liver regeneration in mice is linked to a severe fibrogenic response driven by hepatic progenitor cell activation
  publication-title: Am J Pathol
  doi: 10.1016/j.ajpath.2013.03.018
– volume: 61
  start-page: 2100
  year: 2015
  ident: hep28274-bib-0047-20241017
  article-title: Determining the fate of hepatic cells by lineage tracing: facts and pitfalls
  publication-title: Hepatology
  doi: 10.1002/hep.27659
– volume: 9
  start-page: 231
  year: 2012
  ident: hep28274-bib-0036-20241017
  article-title: The biliary tree—a reservoir of multipotent stem cells
  publication-title: Nat Rev Gastroenterol Hepatol
  doi: 10.1038/nrgastro.2012.23
– volume: 59
  start-page: 1617
  year: 2014
  ident: hep28274-bib-0037-20241017
  article-title: Liver regeneration by stem/progenitor cells
  publication-title: Hepatology
  doi: 10.1002/hep.26753
– volume: 310
  start-page: 68
  year: 2005
  ident: hep28274-bib-0045-20241017
  article-title: Transforming growth factor‐beta signaling in stem cells and cancer
  publication-title: Science
  doi: 10.1126/science.1118389
– volume: 185
  start-page: 325
  year: 2015
  ident: hep28274-bib-0032-20241017
  article-title: A new mdr2−/− mouse model of sclerosing cholangitis with rapid fibrosis progression, early‐onset portal hypertension, and liver cancer
  publication-title: Am J Pathol
  doi: 10.1016/j.ajpath.2014.10.013
– volume: 15
  start-page: 259
  year: 1998
  ident: hep28274-bib-0003-20241017
  article-title: Ductular reaction and its diagnostic significance
  publication-title: Semin Diagn Pathol
– volume: 41
  start-page: 809
  year: 2005
  ident: hep28274-bib-0041-20241017
  article-title: Fibrosis correlates with a ductular reaction in hepatitis C: roles of impaired replication, progenitor cells and steatosis
  publication-title: Hepatology
  doi: 10.1002/hep.20650
– volume: 96
  start-page: 319
  year: 1999
  ident: hep28274-bib-0013-20241017
  article-title: The integrin alphavbeta6 binds and activates latent TGF beta 1: a mechanism for regulating pulmonary inflammation and fibrosis
  publication-title: Cell
  doi: 10.1016/S0092-8674(00)80545-0
– volume: 48
  start-page: 453
  year: 2008
  ident: hep28274-bib-0014-20241017
  article-title: Integrin alphavbeta6 is a marker of the progression of biliary and portal liver fibrosis and a novel target for antifibrotic therapies
  publication-title: J Hepatol
  doi: 10.1016/j.jhep.2007.11.021
– volume: 15
  start-page: 340
  year: 2014
  ident: hep28274-bib-0035-20241017
  article-title: Adult hepatocytes are generated by self‐duplication rather than stem cell differentiation
  publication-title: Cell Stem Cell
  doi: 10.1016/j.stem.2014.06.003
– volume: 13
  start-page: 394
  year: 2008
  ident: hep28274-bib-0028-20241017
  article-title: Regulation of in situ to invasive breast carcinoma transition
  publication-title: Cancer Cell
  doi: 10.1016/j.ccr.2008.03.007
– volume: 31
  start-page: 63
  year: 2013
  ident: hep28274-bib-0022-20241017
  article-title: Mass‐encoded synthetic biomarkers for multiplexed urinary monitoring of disease
  publication-title: Nat Biotechnol
  doi: 10.1038/nbt.2464
– volume: 271
  start-page: 25976
  year: 1996
  ident: hep28274-bib-0043-20241017
  article-title: Identification of a sequence within the integrin beta6 subunit cytoplasmic domain that is required to support the specific effect of alphavbeta6 on proliferation in three‐dimensional culture
  publication-title: J Biol Chem
  doi: 10.1074/jbc.271.42.25976
– volume: 15
  start-page: 605
  year: 2014
  ident: hep28274-bib-0034-20241017
  article-title: Bipotential adult liver progenitors are derived from chronically injured mature hepatocytes
  publication-title: Cell Stem Cell
  doi: 10.1016/j.stem.2014.09.008
– volume: 50
  start-page: 1294
  year: 2009
  ident: hep28274-bib-0005-20241017
  article-title: Targeting liver fibrosis: strategies for development and validation of antifibrotic therapies
  publication-title: Hepatology
  doi: 10.1002/hep.23123
– volume: 44
  start-page: 167
  year: 2006
  ident: hep28274-bib-0050-20241017
  article-title: Oral imatinib treatment reduces early fibrogenesis but does not prevent progression in the long term
  publication-title: J Hepatol
  doi: 10.1016/j.jhep.2005.06.015
– volume: 52
  start-page: 362
  year: 2010
  ident: hep28274-bib-0015-20241017
  article-title: The alphavbeta6 integrin is a highly specific immunohistochemical marker for cholangiocarcinoma
  publication-title: J Hepatol
  doi: 10.1016/j.jhep.2009.12.006
– volume: 61
  start-page: 678
  year: 2015
  ident: hep28274-bib-0019-20241017
  article-title: Connective tissue growth factor and integrin alphavbeta6: a new pair of regulators critical for ductular reaction and biliary fibrosis in mice
  publication-title: Hepatology
  doi: 10.1002/hep.27425
– volume: 133
  start-page: 80
  year: 2007
  ident: hep28274-bib-0042-20241017
  article-title: Progressive fibrosis in nonalcoholic steatohepatitis: association with altered regeneration and a ductular reaction
  publication-title: Gastroenterology
  doi: 10.1053/j.gastro.2007.05.012
– volume: 47
  start-page: 1544
  year: 2008
  ident: hep28274-bib-0006-20241017
  article-title: Clinicopathological study on cholangiolocellular carcinoma suggesting hepatic progenitor cell origin
  publication-title: Hepatology
  doi: 10.1002/hep.22238
– volume: 41
  start-page: 1521
  year: 1993
  ident: hep28274-bib-0011-20241017
  article-title: Restricted distribution of integrin beta 6 mRNA in primate epithelial tissues
  publication-title: J Histochem Cytochem
  doi: 10.1177/41.10.8245410
– volume: 127
  start-page: 1565
  year: 2004
  ident: hep28274-bib-0004-20241017
  article-title: The cholangiopathies: disorders of biliary epithelia
  publication-title: Gastroenterology
  doi: 10.1053/j.gastro.2004.08.006
– volume: 422
  start-page: 169
  year: 2003
  ident: hep28274-bib-0024-20241017
  article-title: Loss of integrin alpha(v)beta6‐mediated TGF‐beta activation causes Mmp12‐dependent emphysema
  publication-title: Nature
  doi: 10.1038/nature01413
– volume: 25
  start-page: 1185
  year: 2011
  ident: hep28274-bib-0048-20241017
  article-title: Foxl1‐Cre‐marked adult hepatic progenitors have clonogenic and bilineage differentiation potential
  publication-title: Genes Dev
  doi: 10.1101/gad.2027811
– volume: 100
  start-page: 11881
  issue: Suppl. 1
  year: 2003
  ident: hep28274-bib-0038-20241017
  article-title: The origin and liver repopulating capacity of murine oval cells
  publication-title: Proc Natl Acad Sci USA
– volume: 51
  start-page: 660
  year: 2009
  ident: hep28274-bib-0001-20241017
  article-title: Diagnosis and management of primary sclerosing cholangitis
  publication-title: Hepatology
– volume: 133
  start-page: 921
  year: 1996
  ident: hep28274-bib-0023-20241017
  article-title: Inactivation of the integrin beta6 subunit gene reveals a role of epithelial integrins in regulating inflammation in the lung and skin
  publication-title: J Cell Biol
  doi: 10.1083/jcb.133.4.921
– volume: 147
  start-page: 1378
  year: 2014
  ident: hep28274-bib-0027-20241017
  article-title: Extrahepatic platelet‐derived growth factor‐beta, delivered by platelets, promotes activation of hepatic stellate cells and biliary fibrosis in mice
  publication-title: Gastroenterology
  doi: 10.1053/j.gastro.2014.08.038
– volume: 136
  start-page: 1951
  year: 2009
  ident: hep28274-bib-0033-20241017
  article-title: Potential hepatic stem cells reside in EpCAM+ cells of normal and injured mouse liver
  publication-title: Development
  doi: 10.1242/dev.031369
– reference: 17924447 - Hepatology. 2007 Nov;46(5):1404-12
– reference: 23354049 - Nature. 2013 Feb 14;494(7436):247-50
– reference: 14722815 - Semin Liver Dis. 2003 Nov;23(4):385-96
– reference: 23242163 - Nat Biotechnol. 2013 Jan;31(1):63-70
– reference: 25203810 - Hepatology. 2015 Feb;61(2):678-91
– reference: 18538673 - Gastroenterology. 2008 Aug;135(2):660-70
– reference: 22371217 - Nat Rev Gastroenterol Hepatol. 2012 Feb 28;9(4):231-40
– reference: 19429791 - Development. 2009 Jun;136(11):1951-60
– reference: 18393293 - Hepatology. 2008 May;47(5):1544-56
– reference: 12634787 - Nature. 2003 Mar 13;422(6928):169-73
– reference: 15521023 - Gastroenterology. 2004 Nov;127(5):1565-77
– reference: 10025398 - Cell. 1999 Feb 5;96(3):319-28
– reference: 20101749 - Hepatology. 2010 Feb;51(2):660-78
– reference: 25503476 - Hepatology. 2015 Jun;61(6):2100-3
– reference: 12902545 - Proc Natl Acad Sci U S A. 2003 Sep 30;100 Suppl 1:11881-8
– reference: 21677751 - Nature. 2011 Jun 15;474(7351):343-9
– reference: 25478810 - Am J Pathol. 2015 Feb;185(2):325-34
– reference: 9845427 - Semin Diagn Pathol. 1998 Nov;15(4):259-69
– reference: 20137822 - J Hepatol. 2010 Mar;52(3):362-9
– reference: 23680654 - Am J Pathol. 2013 Jul;183(1):182-94
– reference: 24681248 - Am J Pathol. 2014 May;184(5):1263-7
– reference: 25130492 - Cell Stem Cell. 2014 Sep 4;15(3):340-9
– reference: 14960589 - J Biol Chem. 2004 Apr 23;279(17 ):17875-87
– reference: 7673344 - J Cell Sci. 1995 Jun;108 ( Pt 6):2241-51
– reference: 17600122 - Am J Pathol. 2007 Aug;171(2):525-36
– reference: 21632826 - Genes Dev. 2011 Jun 1;25(11):1193-203
– reference: 25312494 - Cell Stem Cell. 2014 Nov 6;15(5):605-18
– reference: 19800575 - Cancer Cell. 2009 Oct 6;16(4):295-308
– reference: 8698195 - Gastroenterology. 1996 Jul;111(1):165-71
– reference: 25173753 - Gastroenterology. 2014 Dec;147(6):1378-92
– reference: 8824234 - J Biol Chem. 1996 Oct 18;271(42):25976-80
– reference: 17631134 - Gastroenterology. 2007 Jul;133(1):80-90
– reference: 21632825 - Genes Dev. 2011 Jun 1;25(11):1185-92
– reference: 15793848 - Hepatology. 2005 Apr;41(4):809-18
– reference: 16532004 - Nat Med. 2006 Apr;12(4):410-6
– reference: 16110324 - J Clin Invest. 2005 Sep;115(9):2330-40
– reference: 16879391 - Histopathology. 2006 Aug;49(2):138-51
– reference: 16210527 - Science. 2005 Oct 7;310(5745):68-71
– reference: 15330800 - Histopathology. 2004 Sep;45(3):226-36
– reference: 8245410 - J Histochem Cytochem. 1993 Oct;41(10):1521-7
– reference: 8666675 - J Cell Biol. 1996 May;133(4):921-8
– reference: 18455123 - Cancer Cell. 2008 May;13(5):394-406
– reference: 16214602 - Lancet. 2005 Oct 8;366(9493):1303-14
– reference: 21277850 - Gastroenterology. 2011 May;140(5):1642-52
– reference: 18221819 - J Hepatol. 2008 Mar;48(3):453-64
– reference: 17187411 - Hepatology. 2007 Jan;45(1):31-41
– reference: 24115180 - Hepatology. 2014 Apr;59(4):1617-26
– reference: 16168515 - J Hepatol. 2006 Jan;44(1):167-75
– reference: 16223543 - J Hepatol. 2005 Dec;43(6):1045-54
– reference: 19711424 - Hepatology. 2009 Oct;50(4):1294-306
SSID ssj0009428
Score 2.5095665
Snippet Integrin αvβ6 is rapidly up‐regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent...
Integrin αvβ6 is rapidly up-regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent...
UNLABELLEDIntegrin αvβ6 is rapidly up-regulated on cells of epithelial lineage during tissue injury, where one of its primary functions is activation of latent...
SourceID pubmedcentral
proquest
pubmed
crossref
wiley
SourceType Open Access Repository
Aggregation Database
Index Database
Enrichment Source
Publisher
StartPage 217
SubjectTerms Animals
Antigens, Neoplasm - physiology
Carcinogenesis
Cholangitis, Sclerosing - etiology
Fibrosis - etiology
Hepatocytes
Humans
Integrins - physiology
Liver - pathology
Liver Cirrhosis - etiology
Liver Neoplasms - etiology
Male
Mice
Mice, Inbred C57BL
Stem Cells - physiology
Title Integrin αvβ6 critically regulates hepatic progenitor cell function and promotes ductular reaction, fibrosis, and tumorigenesis
URI https://onlinelibrary.wiley.com/doi/abs/10.1002%2Fhep.28274
https://www.ncbi.nlm.nih.gov/pubmed/26448099
https://www.proquest.com/docview/1751488536
https://pubmed.ncbi.nlm.nih.gov/PMC5312042
Volume 63
hasFullText 1
inHoldings 1
isFullTextHit
isPrint
journalDatabaseRights – providerCode: PRVWIB
  databaseName: Wiley Online Library Full Collection 2020
  customDbUrl:
  eissn: 1527-3350
  dateEnd: 99991231
  omitProxy: false
  ssIdentifier: ssj0009428
  issn: 0270-9139
  databaseCode: DRFUL
  dateStart: 19960101
  isFulltext: true
  titleUrlDefault: https://onlinelibrary.wiley.com
  providerName: Wiley-Blackwell
link http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwpV3dS9xAEB_0rhRf-mWtV1tZpQ8-GI3JJpvQp9J6WLDHIVruLexmdzFgc3K5E_ron2T_EP8mZza5XA9bKBTyEMhk2Y-Z2d_sx28APkRGC62E9Hia-B6XnHsKgbKH6qF0Elubu1yH30_FYJCMRulwBT7O78LU_BDtghtZhvPXZOBSVYcL0tBLc32A8YLgq9ANUG95B7pfzvoXpwvOXe5Sq2Lg5dMGczonFvKDw_bn5enoEcZ8fFTydwjr5qD-8_-q_Qt41kBP9qnWlZewYspX8PRbs7m-DrdfHXVEUbL7u5v7XzHLmzQIVz_ZpE5ZbyqGRRLJK6ODXYb8wYTR4j-jCZIGmclS00dUAZQmNlk66IoF1Dco9pnFeo-rotp3ktPZD8rNRR63qF7DRf_4_POJ12Ro8HKyfS_MA41GjZhHSx0qnOdyG8lQoReNQi4pnbofWS6t4PgcWYXhkhaxQsxhtbIm3IBOOS7NJjAVHkkVpYlBDeE2tlIk1uSh9rVSRqa8B3vzgcryhr6csmhcZTXxcpBh-zPXpT3YbUWva86OPwntzEc7Q4uinpKlGc-qDAEVxojYgLgHb-rRb4tx4SyC6h6IJb1oBYite_lLWVw61m50dgF6SGyH04u_1yw7OR66l7f_LroFa4jkmrWhd9CZTmbmPTzJb6ZFNdmGVTFKthvzeACtnBiU
linkProvider Wiley-Blackwell
linkToHtml http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwpV1LT9tAEB7RgNpeCm0pTQtlqTj0EIOJ1y-plwqIghoihKDiZu16d4UlcKI4idRjfxL8EH5TZ9aOIQKkSpV8sOTxah8zs9_s4xuAbV-rUMlQODyOXIcLzh2JQNlB9ZAqCoxJba7DX72w348uLuKTBfg-uwtT8kPUC25kGdZfk4HTgvTuPWvopR7uYMAQ8hewyFGN_AYsHpx2znv3pLvc5lbFyMulHeZ4xizktnfrn-fno0cg8_FZyYcY1k5CneX_q_4KvKnAJ_tRastbWND5O3h5XG2vv4c_R5Y8IsvZ3c307jZgaZUI4eo3G5VJ63XBsEiieWV0tEuTRxgxWv5nNEXSMDORK_qISoDSxCdLR12xgPIORYsZrPigyIqWlRxPrik7F_ncrFiF887h2X7XqXI0OClZv-OlbYVmjahHCeVJnOlS4wtPoh_1PS4oobrrGy5MyPHZMxIDJhUGElGHUdJo7wM08kGuPwKT3p6Qfhxp1BFuAiPCyOjUU66SUouYN-HbbKSStCIwpzwaV0lJvdxOsP2J7dImfK1FhyVrx1NCW7PhTtCmqKdErgeTIkFIhVEiNiBowlo5_HUxNqBFWN2EcE4xagHi657_kmeXlrcb3V0bfSS2wyrG8zVLuocn9uXTv4tuwqvu2XEv6R31f36G14jrqpWidWiMRxO9AUvpdJwVoy-VlfwFvP4bnA
linkToPdf http://cvtisr.summon.serialssolutions.com/2.0.0/link/0/eLvHCXMwpV1LSxxBEC6MiuRi1ERd46OVHHJwdNzpeYEXURfFdVkkBm9D93Q3DpjZZWdXyDE_yfwQf1OqemZHFw0IwhwGpqbpR1X1V_34CuCbr1WoZCgcHkeuwwXnjkSg7KB6SBUFxqQ21-HPdtjpRDc3cXcKDsd3YUp-iHrBjSzD-msycN1XZv-JNfRW9_cwYAj5B5jhfhygWc6cXLWu20-ku9zmVsXIy6Ud5njMLOQ29-ufJ-ejFyDz5VnJ5xjWTkKtT--r_gLMV-CTHZXasghTOl-Cuctqe_0z_Dm35BFZzh4f7h__BiytEiHc_WaDMmm9LhgWSTSvjI52afIIA0bL_4ymSBpmJnJFH1EJUJr4ZOmoKxZQ3qHYZQYr3iuyYtdKDke_KDsX-dys-ALXrdMfx2dOlaPBScn6HS9tKjRrRD1KKE_iTJcaX3gS_ajvcUEJ1V3fcGFCjs-BkRgwqTCQiDqMkkZ7yzCd93K9Ckx6B0L6caRRR7gJjAgjo1NPuUpKLWLegO_jkUrSisCc8mjcJSX1cjPB9ie2SxuwU4v2S9aO14S2x8OdoE1RT4lc90ZFgpAKo0RsQNCAlXL462JsQIuwugHhhGLUAsTXPfklz24tbze6uyb6SGyHVYz_1yw5O-3al7W3i27BXPeklbTPOxdf4SPCumqhaB2mh4OR3oDZ9H6YFYPNykj-ASQXGxc
openUrl ctx_ver=Z39.88-2004&ctx_enc=info%3Aofi%2Fenc%3AUTF-8&rfr_id=info%3Asid%2Fsummon.serialssolutions.com&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Integrin+%CE%B1v%CE%B26+critically+regulates+hepatic+progenitor+cell+function+and+promotes+ductular+reaction%2C+fibrosis%2C+and+tumorigenesis&rft.jtitle=Hepatology+%28Baltimore%2C+Md.%29&rft.au=Peng%2C+Zhen%E2%80%90Wei&rft.au=Ikenaga%2C+Naoki&rft.au=Liu%2C+Susan+B.&rft.au=Sverdlov%2C+Deanna+Y.&rft.date=2016-01-01&rft.issn=0270-9139&rft.eissn=1527-3350&rft.volume=63&rft.issue=1&rft.spage=217&rft.epage=232&rft_id=info:doi/10.1002%2Fhep.28274&rft.externalDBID=n%2Fa&rft.externalDocID=10_1002_hep_28274
thumbnail_l http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/lc.gif&issn=0270-9139&client=summon
thumbnail_m http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/mc.gif&issn=0270-9139&client=summon
thumbnail_s http://covers-cdn.summon.serialssolutions.com/index.aspx?isbn=/sc.gif&issn=0270-9139&client=summon